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Leptobarbus Bleeker 1859    leptos, thin, referring to slender shape of L. hoevenii; Barbus, referring to previous placement of L. hoevenii in that genus

Leptobarbus hoevenii (Bleeker 1851)    in honor of Bleeker’s fellow Dutchman, “le célèbre professeur de zoologie” Jan van der Hoeven (1801-1868)

Leptobarbus hosii (Regan 1906)    in honor of Charles Hose (1863-1929), British colonial administrator in Borneo, zoologist and ethnologist, who collected type

Leptobarbus melanopterus Weber & de Beaufort 1916    melanos, black; pterus, fin, referring to nearly all-black ventral, pectoral and dorsal fins

Leptobarbus melanotaenia Boulenger 1894    melanos, black; taenia, band, referring to black lateral stripe running along body above lateral line and/or black vertical bar on posterior border of gill opening

Leptobarbus rubripinna (Fowler 1937)    ruber, red; pinna, fin, referring to “more or less bright orange red” coloration of fins (except dorsal) in living specimens

Subfamily ACHEILOGNATHINAE Bitterlings
6 genera • 83 species/subspecies

Acheilognathus Bleeker 1859    a-, without; cheilos, lip; gnathos, jaw, referring to lower jaw without (or with only a lateral) labial fold

Acheilognathus asmussii (Dybowski 1872)    patronym not identified, possibly in honor of German entomologist Eduard Assmuss (1838-1882)

Acheilognathus barbatulus Günther 1873    diminutive of barbatus, bearded, i.e., a small beard, referring to “pair of very short barbels”

Acheilognathus barbatus Nichols 1926    bearded, referring to well-developed terminal barbel (compared to the barbelless A. gracilis, described in same paper)

Acheilognathus binidentatus Li 2001    binus, two by two; dentatus, toothed, but Li erroneously says name means “two spines,” referring to two unbranched rays (not spines) on pelvic fin

Acheilognathus brevicaudatus Chen & Li 1987    brevis, short; caudatus, tailed, referring to shorter caudal peduncle compared to A. elongatus

Acheilognathus changtingensis Yang, Zhu, Xiong & Liu 2011    ensis, suffix denoting place: Changting County, Fujian Province, China, where it appears to be endemic

Acheilognathus chankaensis (Dybowski 1872)    ensis, suffix denoting place: Lake Khanka, southeastern Russia, type locality

Acheiloganthus coreanus Steindachner 1892    Korean, referring to type locality in Seoul, Korea

Acheilognathus cyanostigma Jordan & Fowler 1903    cyano-, dark blue; stigma, mark, referring to blue-black lateral stripe

Acheilognathus deignani (Smith 1945)    in honor of Herbert Girton Deignan (1906-1968), Associate Curator of Birds, U.S. National Museum, who collected type

Acheilognathus elongatoides Kottelat 2001    oides, having the form of: replacement name for A. elongatus Mai 1978 (preoccupied by A. elongatus [Regan 1908]), probably referring to long, compressed body, longer than morphologically similar Pararhodeus (=Rhodeus) fangi

Acheilognathus elongatus (Regan 1908)    referring to more elongate body compared to A. atranalis (=Acanthorhodeus chankaensis)

Acheilognathus fasciodorsalis Nguyen 2001    fascio-, band; dorsalis, dorsal, referring to three black stripes on dorsal fin

Acheilognathus gracilis Nichols 1926    thin or slender, probably referring to body shape compared to A. barbatus (described in the same paper), which has an elevated nape

Acheilognathus hypselonotus (Bleeker 1871)    hypselo-, high; notus, back, referring to convex body shape (common to many congeners)

Acheilognathus imberbis Günther 1868    beardless, referring to absence of barbels

Acheilognathus imfasciodorsalis Nguyen 2001    im-, not; fascio-, band; dorsalis, dorsal, lacking three black stripes on dorsal fin as seen on A. fasciodorsalis

Acheilognathus kyphus (Mai 1978)    referring to Ky Phu stream in Bac Thai province (Thailand), presumably type locality or close to it

Acheilognathus lanchiensis (Herre & Lin 1936)    ensis, suffix denoting place: Lanchie, upper Tsien Tang River, Chekiang Province, China, type locality

Acheilognathus longibarbatus (Mai 1978)    longus, long; barbatus, bearded, referring to long maxillary barbel, equal to or longer than eye diameter

Acheilognathus longipinnis Regan 1905    longus, long; pinnis, fin, referring to long pectoral fin, nearly as long as head, extending to root of ventral

Acheilognathus macromandibularis Doi, Arai & Liu 1999    macro-, large; mandibularis, of the jaw, referring to large mouth, reaching posteriorly to a vertical line through anterior margin of orbit

Acheilognathus macropterus (Bleeker 1871)    macro-, long; pterus, fin, probably referring to long and high dorsal fin, base of which is ~3 times longer than base of tail

Acheilognathus majusculus Kim & Yang 1998    somewhat bigger, referring to larger body size compared to other bitterlings

Acheilognathus melanogaster Bleeker 1860    melano-, black; gaster, belly, referring to black abdomen

Acheilognathus meridianus (Wu 1939)    anus, belonging to: meridies, south, referring to distribution in southern China

Acheilognathus microphysa Yang, Chu & Chen 1990    micro-, small; physis, air bladder, referring to greatly reduced posterior chamber of air bladder

Acheilognathus nanchongensis Deng 1996    ensis, suffix denoting place: Nanchong, Sichuan Province, China, where it appears to be endemic

Acheilognathus nguyenvanhaoi Nguyen, Tram & Ta 2013    in honor of Vietnamese ichthyologist Nguyen Van Hao

Acheilognathus omeiensis (Shih & Tchang 1934)    ensis, suffix denoting place: Omei, Sichuan Province, China, type locality

Acheilognathus peihoensis (Fowler 1910)    ensis, suffix denoting place: Pei-Ho River, at Tien Tsin (now Tianjin), former Pechili Province, China, type locality

Acheilognathus polylepis (Wu 1964)    poly, many; lepis, scales, having more lateral line scales than A. barbatulus and A. tonkinensis

Acheilognathus polyspinus (Holcík 1972)    poly, many; spinus, spine, referring to large number of rays in dorsal and anal fins

Acheilognathus rhombeus (Temminck & Schlegel 1846)    presumably referring to rhomboid body shape

Acheilognathus striatus Yang, Xiong, Tang & Liu 2010    stripe, referring to broad longitudinal stripe on body

Acheilognathus tabira tabira Jordan & Thompson 1914    vernacular for this species at Lake Biwa, Japan

Acheilognathus tabira erythropterus Arai, Fujikawa & Nagata 2007    erythro, red; pterus, fin, referring to anal fin edged with red in nuptial males

Acheilognathus tabira jordani Arai, Fujikawa & Nagata 2007    in honor of David Starr Jordan (1851-1931), who contributed greatly to Japanese ichthyology, including original description of A. tabira

Acheilognathus tabira nakamurae Arai, Fujikawa & Nagata 2007    in honor of Morizumi Nakamura (1914-1998), for contributions to the systematics of Japanese bitterlings

Acheilognathus tabira tohokuensis Arai, Fujikawa & Nagata 2007    ensis, suffix denoting place: Tohoku area of Japan, where it is distributed

Acheilognathus taenianalis (Günther 1873)    taenia, band; analis, anal, probably referring to two rows of white spots (“particularly distinct” in males) and/or white margin on anal fin

Acheilognathus tonkinensis (Vaillant 1892)    ensis, suffix denoting place: upper Tonkin, northern Viêt Nam, type locality

Acheilognathus typus (Bleeker 1863)    being the type species of Pseudoperilampus, now a junior synonym of Rhodeus

Acheilognathus yamatsutae Mori 1928    in honor of Mr. K. Yamatsuta, a teacher at the Mukden Higher Girls School, who “obtained … a fine type specimen”

Paratanakia Chang, Chen & Mayden 2014    para-, near, referring to morphological similarity of P. himantegus to Tanakia

Paratanakia himantegus himantegus (Günther 1868)    himantos, strap or leash; egos, I lead, i.e., handling or leading with a strap or leash, probably referring to long ovipositor of breeding females

Paratanakia himantegus chii (Miao 1934)    in honor of Chen-Ju Ch’i, Director of Bureau of Education of Honan [now Henan] Province, China, for his “kind support” of the author’s study of Kiangsu fishes

Pseudorhodeus Chang, Chen & Mayden 2014    pseudo-, false, referring to similarity (e.g., incomplete lateral line) but not close relationship to Rhodeus

Pseudorhodeus tanago (Tanaka 1909)    Japanese vernacular for bitterling, translating as “fishes on palm”

Rhodeus Agassiz 1832    rose, referring to rosy color of breeding males

Rhodeus albomarginatus Li & Arai 2014    albus, white; marginatus, margined, referring to diagnostic white margin on anal fin of adult male

Rhodeus amarus (Bloch 1782)   bitter, referring to taste of its roe

Rhodeus amurensis (Vronsky 1967)    ensis, suffix denoting place: lower Amur River drainage, Russia, type locality

Rhodeus atremius (Jordan & Thompson 1914)   a-, without; tremia, aperture, referring to “completely absent” lateral line

Rhodeus colchicus Bogutskaya & Komlev 2001    icus, belonging to: Colchis, ancient name for the eastern coast of the Black Sea in Georgia (Eurasia), type locality

Rhodeus fangi (Miao 1934)   in honor of P. W. Fang, Metropolitan Museum of Natural History, Academia Sinica (spelled “Sinerica” by Miao)

Rhodeus haradai Arai, Suzuki & Shen 1990    in honor of I. Harada, ichthyologist who identified this new species as R. spinalis in 1943

Rhodeus hondae (Jordan & Metz 1913)    in honor of K. Honda, director, Agricultural Station at Suigen (Korea), “who obtained for us a fine collection from the pond at this station”

Rhodeus laoensis Kottelat, Doi & Musikasinthorn 1998    ensis, suffix denoting place: Laos, where it is endemic

Rhodeus lighti (Wu 1931)    in honor of zoologist Sol Felty Light (1886-1947), University of California, for “constant help and encouragement” in Wu’s zoological studies during their three-year association at University of Amoy (now Xiamen University)

Rhodeus mantschuricus Mori 1934    Manchurian, referring to distribution in northeast Asia (Amur River basin, Russia and China)

Rhodeus meridionalis Karaman 1924    southern, allusion not explained, perhaps referring to Lake Dojran, type locality, in southern Macedonia at border with Greece

Rhodeus monguonensis (Li 1989)    ensis, suffix denoting place: Mongolia, referring to distribution in Inner Mongolia Autonomous Region

Rhodeus notatus Nichols 1929    marked, probably referring to dark markings on caudal peduncle, middle caudal rays, behind upper angle of gill cleft, snout, and upper part of gill cover

Rhodeus ocellatus ocellatus (Kner 1866)    eyespot, referring to black spot behind shoulder

Rhodeus ocellatus kurumeus Jordan & Thompson 1914    eus, adjectival suffix: Kurume, Kiusiu, Japan, type locality

Rhodeus ocellatus vietnamensis Mai 1978    ensis, suffix denoting place: North Viêt Nam, where it is endemic

Rhodeus pseudosericeus Arai, Jeon & Ueda 2001    pseudo-, false, i.e., although similar to R. sericeus, such an appearance is false

Rhodeus rheinardti (Tirant 1883)    patronym not explained, possibly in honor of M. le commandant Rheinhart (note spelling), a French official in Hué, Viêt Nam (type locality for this bitterling), who sent zoological specimens to Paris

Rhodeus sciosemus (Jordan & Thompson 1914)    scio-, shade or shadow; sema, mark, probably referring to large black spot on dorsal fin (absent on R. atremius, described in same paper)

Rhodeus sericeus (Pallas 1776)    silken, perhaps referring to soft, lustrous color of breeding males

Rhodeus shitaiensis Li & Arai 2011    ensis, suffix denoting place: Shitai County, Anhui Province, China, type locality

Rhodeus sinensis Günther 1868    ensis, suffix denoting place: Sinica (China), where it is endemic

Rhodeus smithii (Regan 1908)    in honor of British traveler, sportsman and naturalist Richard Gordon Smith (1858-1918), who collected specimens in Japan for the British Museum (Natural History), including type of this one

Rhodeus spinalis Oshima 1926    spiny, referring to osseus dorsal and anal fin spines

Rhodeus suigensis (Mori 1935)    ensis, suffix denoting place: Suigen, Korea, type locality

Rhodeus uyekii (Mori 1935)    in honor of Homiki Uyeki (1882-?), botanist, Suigen Agricultural College (type locality is in Suigen, Korea)

Sinorhodeus Li, Liao & Arai 2017    Sino-, Chinese, referring to distribution in China; Rhodeus, referring to morphological similarity with that genus

Sinorhodeus microlepis Li, Liao & Arai 2017    micro-, small; lepis, scale, referring to smaller and therefore more numerous scales compared to any species in Rhodeus

Tanakia Jordan & Thompson 1914    ia, belonging to: Shigeho Tanaka (1878-1974), “accomplished” ichthyologist of the Imperial University of Tokyo, who described T. shimazui in 1908 and Pseudorhodeus tanago in 1909

Tanakia intermedia (Temminck & Schlegel 1846)    ensis, suffix denoting place: South Korea, where it is endemic

Tanakia koreensis (Kim & Kim 1990)    ensis, suffix denoting place: South Korea, where it is endemic

Tanakia lanceolata (Temminck & Schlegel 1846)    presumably referring to its more lanceolate (leaf-like) shape compared to other Japanese bitterlings

Tanakia latimarginata Kim, Jeon & Suk 2014    latus, broad; marginata, margined, referring to broad black border on anal fin of mature males

Tanakia limbata (Temminck & Schlegel 1846)    bordered, presumably referring to blackish edges on dorsal and anal fins

Tanakia shimazui (Tanaka 1908)    in honor of “Mr. Shimazu” (no forename given), a naturalist in Tokyo, who collected type

Tanakia signifer (Berg 1907)    signum, mark; fero, to bear, probably referring to broad whitish band on dorsal fin margin

Tanakia somjinensis (Kim & Kim 1991)    ensis, suffix denoting place: Somjin River, South Korea, type locality

Subfamily GOBIONINAE Gudgeons
30 genera/subgenera – 232 species/subspecies

Abbottina Jordan & Fowler 1903    –ina, belonging to: James Francis Abbott, professor, Japanese Military Academy at Etajima, who assisted David Starr Jordan and John Otterbein Snyder in their work collecting fishes in Japan        

Abbottina binhi Nguyen 2001    in honor of Bình Nguyén, who helped Nguyen collect type (Ngo S. V., pers. comm.)

Abbottina lalinensis Huang & Li 1995    ensis, suffix denoting place: Lalin, Heilongjiang Province, China, type locality

Abbottina liaoningensis Qin 1987    ensis, suffix denoting place: Liaoning Province, China, type locality   

Abbottina obtusirostris (Wu & Wang 1931)    obtusus, blunt; rostris, snout, referring to blunt snout, “without sudden incision” 

Abbottina rivularis (Basilewsky 1855)    of a small stream or brook, “occurring in the pools of a brook” (translation)           

Acanthogobio Herzenstein 1892    acanthus, thorn or spine, referring to strong osseus spine on dorsal fin, i.e., a spiny or thorny Gobio                 

Acanthogobio guentheri Herzenstein 1892    in honor of ichthyologist and herpetologist Albert Günther (1830-1914)            

Belligobio Jordan & Hubbs 1925    etymology not explained, perhaps bellus, beautiful, referring to “striking” coloration of type species, B. eristigma (= Hemibarbus mylodon), i.e., a “beautiful Gobio”             

Belligobio nummifer (Boulenger 1901)    nummus, coin; fero, to bear, referring to series of six round (i.e., coin-shaped) black spots along body and tail, above lateral line           

Belligobio pengxianensis Luo, Le & Chen 1977    ensis, suffix denoting place: Pengxian County, Sichuan Province, China, type locality      

Biwia Jordan & Fowler 1903    ia, belonging to: Lake Biwa, distribution of type species, B. zezera              

Biwia springeri (Bănărescu & Nalbant 1973)    in honor of ichthyologist Victor G. Springer (b. 1928), U.S. National Museum, who collected type  

Biwia tama Oshima 1957    named for Tama River, Denyenchofu, Japan, where it is endemic       

Biwia yodoensis Kawase & Hosoya 2010    ensis, suffix denoting place: Yodo River basin (including Lake Biwa), Japan, where it is endemic     

Biwia zezera (Ishikawa 1895)    Japanese name of a small minnow from Zeze, a village near Lake Biwa, type locality           

Coreius Jordan & Starks 1905    Korean, presumably referring to two specimens of C. cetopsis collected at Chemulpo, Korea (also occurs in China)    

Coreius cetopsis (Kner 1867)    cetos, whale; opsis, appearance, allusion not explained nor evident

Coreius guichenoti (Sauvage & Dabry de Thiersant 1874)    patronym not identified but clearly in honor of the authors’ colleague, zoologist Antoine Alphone Guichenot (1809-1876), Muséum national d’Histoire naturelle (Paris) 

Coreius heterodon (Bleeker 1864)    heteros, different; odon, tooth, having one row of raptatorial (hooked inward at the tips) pharyngeal teeth instead of two    

Coreius septentrionalis (Nichols 1925)    northern, probably referring to distribution in northern China (e.g., Mongolia) 

Coreoleuciscus Mori 1935    Corea, alternate spelling of Korea, referring to country where genus is endemic, presumed to be “closely related” to Leuciscus and allied genera (i.e, the Korean Leuciscus)            

Coreoleuciscus aeruginos Song & Bang 2015    Latin for verdigris, referring to bluish-green coloration

Coreoleuciscus splendidus Mori 1935    bright or shining, presumably referring to splendid coloration: sides with two longitudinal bands (one bluish, the other yellowish), and yellowish fins with two (dorsal and caudal) or one crossbars 

Gnathopogon Bleeker 1860    gnathos, jaw; pogon, beard, referring to pair of barbels only on upper lip (or jaw)              

Gnathopogon caerulescens (Sauvage 1883)    bluish, referring to broad blue band along lateral line            

Gnathopogon elongatus elongatus (Temminck & Schlegel 1846)    referring to elongate, compressed body 

Gnathopogon elongatus suwae Jordan & Hubbs 1925    of Lake Suwa at Kamisuwa, Shinshu, Japan, where it is endemic                  

Gnathopogon herzensteini (Günther 1896)    patronym not identified but clearly in honor of Russian ichthyologist Solomon Markovich Herzenstein (1854-1894), who named an Acanthogobio after Günther in 1892   

Gnathopogon imberbis imberbis (Sauvage & Dabry de Thiersant 1874)    beardless, referring to absence of barbels (compared to other species of Gobio, genus in which it was described)                  

Gnathopogon imberbis taeniatus (Günther 1896)    banded, referring to blackish band from upper part of gill opening to root of caudal fin              

Gnathopogon nicholsi (Fang 1943)    in honor of John Treadwell Nichols (1883-1958), curator of fishes at the American Museum of Natural History, who described this species in 1925 but used a preoccupied name, Leucogobio imberbis

Gnathopogon polytaenia (Nichols 1925)    poly, many; taenia, band, probably referring to “indistinct pale streaks along back” and/or “dark and silvery streaks” below lateral line              

Gnathopogon strigatus (Regan 1908)    furrowed or grooved, presumably referring to dark longitudinal stripes along body, which appear to form a furrow between rows of scales       

Gnathopogon taeniellus (Nichols 1925)    taenia, band; –ellus, a diminutive, allusion not explained, possibly referring to fewer, smaller and/or paler streaks on body compared to G. polytaenia           

Gnathopogon tsinanensis (Mori 1928)    ensis, suffix denoting place: Tsi-nan (now Jinan), Shandong Province, China, type locality                

Gobio Cuvier 1816    tautonymous with Cyprinus gobio, derived from the Latin gobionem, from the Greek kobios, a kind of fish, usually applied to gudgeons and gudgeon-like fishes                 

Gobio acutipinnatus Men’shikov 1939    acutus, sharp or pointed; pinnis, fin, referring to more pointed pectoral and ventral fins compared to G. g. gobio    

Gobio alverniae Kottelat & Persat 2005    of Alvernia, a Roman province inhabited by the Arverne tribe, approximately corresponding to the Auvergne region of France (Dept. Haute-Lorie, type locality)   

Gobio artvinicus Turan, Japoshvili, Aksu & Bektaş 2016    –icus, belonging to: Artvin, Turkey, city and eponymous province, type locality

Gobio baliki Turan, Kaya, Bayçelebi, Aksu & Bektaş 2017    in honor of Süleyman Balik, for his contributions to the knowledge of Turkish fishes

Gobio battalgilae Naseka, Erk’akan & Küçük 2006    in honor of ichthyologist Fahire Battalgil (later Battalgazi, 1902-1948), for her contributions to the knowledge of Turkish freshwater fishes

Gobio brevicirris Fowler 1976    brevis, short; cirrus, curl or tendril, referring to shorter barbels (reaching to, not beyond, anterior eye margin) compared to Black and Caspian Sea basin congeners (name coined by Berg in 1914 but not validly published)              

Gobio bulgaricus Drensky 1926    Bulgarian, referring to type locality in Maritsa (also occurs in Turkey, Greece and Macedonia)         

Gobio carpathicus Vladykov 1925    Carpathian, referring to distribution in Carpathian Mountain region of Central Europe  

Gobio coriparoides Nichols 1925    oides, having the form of: genus Coripareius (= Coreius), into which Nichols was “tempted to place this species … were its teeth not typical of Gobio”    

Gobio cynocephalus Dybowski 1869    cyno-, dog; cephalus, head, probably referring to long snout   

Gobio delyamurei Freyhof & Naseka 2005    in honor of Semion Lyudvigovich Delyamure, author of several papers and books on Crimean fishes                  

Gobio feraeensis Stephanidis 1973    ensis, suffix denoting place: Ferae, ancient town that once stood at type locality, ~25 km away from Velestinon, Greece                  

Gobio fushunensis Xie, Li & Xie 2007    ensis, suffix denoting place: Fushun, Liaoning Province, China, where type locality (Hun He [“Muddy”] River is situated

Gobio gobio gobio (Linnaeus 1758)    derived from the Latin gobionem, from the Greek kobios, a kind of fish, usually applied to gudgeons and gudgeon-like fishes              

Gobio gobio chipingi Bănărescu & Nalbant 1964    in honor of Prof. Chi-Ping, who collected type               

Gobio gymnostethus Ladiges 1960    gymno, bare or naked; stethos, breast or chest, referring to “mostly” (translation) scaleless throat            

Gobio hettitorum Ladiges 1960    etymology not explained but probably referring to the Hittites (Latin: Hetthaeis), an ancient Anatolian culture (~1750-1180 BC), presumably referring to distribution in south-central Turkey   

Gobio holurus Fowler 1976    holo-, entire; ourus, tail, referring to less emarginated (slightly forked) caudal fin compared to G. lepidolaemus      

Gobio huanghensis Luo, Le & Chen 1977    ensis, suffix denoting place: Huang He (Yellow River), which runs through Lanzhou, Gansu Province, China, type locality        

Gobio insuyanus Ladiges 1960    anus, belonging to: Insuyu Creek, Cihanbeyli, Turkey, type locality  

Gobio intermedius Battalgil 1943    intermediate in number of throat scales (20-22) between neck and anterior base of back compared to G. gobio (14-15) and G. microlepidotus (30-35)  

Gobio kizilirmakensis Turan, Japoshvili, Aksu & Bektaş 2016    –ensis, suffix denoting place: Kızılırmak River basin, Turkey, where it is endemic

Gobio kovatschevi Chichkoff 1937    in honor of the late Vasily Kovacev, for his research of Bulgarian fishes and especially for his book Freshwater Fish Fauna of Bulgaria (1922)   

Gobio krymensis Bănărescu & Nalbant 1973    ensis, suffix denoting place: southern Crimea, Ukraine, where it is endemic       

Gobio kubanicus Vasil’eva 2004    icus, belonging to: Kuban River drainage, Russia, where it is endemic                 

Gobio lepidolaemus lepidolaemus Kessler 1872    lepido-, scale; laimos, throat, referring to usually scaled throat (naked anteriorly in some specimens)     

Gobio lepidolaemus caucasicus Kamensky 1901    icus, belonging to: northern Caucasus region, Russia, type locality               

Gobio lingyuanensis Mori 1934    ensis, suffix denoting place: Lingyuan, Liaoning Province, China, type locality

Gobio lozanoi Doadrio & Madeira 2004    in honor of Luis Lozano Rey (1878-1958), University of Madrid, for his contribution to the knowledge of Iberian freshwater fishes             

Gobio macrocephalus Mori 1930    macro-, long; cephalus, head, referring to longer head than G. gobio        

Gobio maeandricus Naseka, Erk’akan & Küçük 2006    icus, belonging to: Great Menderes River at Likli, Turkey, type locality             

Gobio meridionalis Xu 1987    southern, referring to distribution south of Yellow River in Henan Province, China            

Gobio microlepidotus Battalgil 1942    micro-, small; lepidotus, scaled, referring to small throat scales, 30-35 between neck and anterior base of back     

Gobio nigrescens (Keyserling 1861)    blackish, presumably referring to 6-7 irregularly shaped, black or brown blotches on back behind dorsal-fin base

Gobio obtusirostris Valenciennes 1842    obtusus, blunt; rostris, snout, “characterized by short muzzle, fat and obtuse” (translation)  

Gobio occitaniae Kottelat & Persat 2005    of Occitanie, the southern part of France, which encompasses most of the range of this species          

Gobio ohridanus Karaman 1924    –anus, belonging to: Lake Ohrid, Republic of Macedonia, type locality (also occurs in Mat River, Albania)     

Gobio rivuloides Nichols 1925    oides, having the form of: presumably the cyprinidontiform genus Rivulus, perhaps referring to flat-topped head

Gobio sakaryaensis Turan, Ekmekçi, Luskova & Mendel 2012    ensis, suffix denoting place: Sakarya River drainage (northwest Anatolia, Black Sea basin), Turkey, where it is endemic                  

Gobio sarmaticus Berg 1949    icus, belonging to: Sarmatia, ancient name for territory corresponding to modern southern Russia, the eastern Balkans, and the distribution of this species in the Ukraine 

Gobio sibiricus Nikolskii 1936    Siberian, referring to distribution in western Siberia (Russia, Kazakhstan and Mongolia)  

Gobio skadarensis Karaman 1937    ensis, suffix denoting place: Skadar Lake (Montenegro and Albania), type locality (but occurs throughout Skadar Lake basin)   

Gobio soldatovi Berg 1914    in honor of ichthyologist Vladimir Konstantinovich Soldatov (1875-1941), who collected type           

Gobio tchangi Li 2015    in honor of ichthyologist Tchunlin (or Tchung-Lin) Tchang (1897-1963)

Gobio volgensis Vasil’eva, Mendel, Vasil’ev, Lusk & Lusková 2008    ensis, suffix denoting place: Volga River basin, Russia, where it appears to be endemic      

Gobiobotia Kreyenberg 1911    gobio, gudgeon; Botia, an Asian loach, likely reflecting Kreyenberg’s provisional placement of G. pappenheimi in the loach family Cobitidae

Subgenus Gobiobotia

Gobiobotia brevibarba Mori 1935    brevis, short; barbus, barbel, referring to eight short barbels, shorter than those on G. pappenheimi       

Gobiobotia brevirostris Chen & Cao 1977    brevis, short; rostris, snout, referring to shorter snout compared to G. pappenheimi and G. tungi

Gobiobotia cheni Bănărescu & Nalbant 1966    in honor of Johnson T. F. Chen, Director of the Taiwan Museum and professor, National Taiwan University, who provided type specimens                 

Gobiobotia filifer (Garman 1912)    filum, thread; fero, to bear, presumably referring to maxillary barbels, which extend beyond eye           

Gobiobotia guilingensis Chen 1989    ensis, suffix denoting place: Guilin, Guangxi Province, China, type locality        

Gobiobotia homalopteroidea Rendahl 1932    oides, having the form of: referring to general resemblance to loach genus Homaloptera (Balitoridae) 

Gobiobotia jiangxiensis Zhang & Liu 1995    ensis, suffix denoting place: Jiangxi Province, China, where it is endemic            

Gobiobotia kolleri Bănărescu & Nalbant 1966    in honor of the late Otto Koller, first ichthyologist to examine specimens on which species is based (1927)        

Gobiobotia longibarba Fang & Wang 1931    longus, long; barba, barbel, referring to long barbels, which extend beyond base of pectoral fin            

Gobiobotia macrocephala Mori 1935    macro-, large; cephalus, head, referring to large head, 3.75 in body length             

Gobiobotia meridionalis Chen & Cao 1977    southern, described as a southern subspecies of G. longibarba

Gobiobotia naktongensis Mori 1935    ensis, suffix denoting place: location not specified, presumably from Naktong (also spelled Nakdong) River basin, South Korea    

Gobiobotia nicholsi Bănărescu & Nalbant 1966    in honor of John Treadwell Nichols (1883-1958), curator of fishes at the American Museum of Natural History, who recognized this species as G. pappenheimi in 1928

Gobiobotia pappenheimi Kreyenberg 1911    patronym not identified but clearly in honor of  Paul Pappenheim (1878-1945), curator of fishes, Royal Museum of Berlin, who authored an addendum to the description reflecting Kreyenberg’s uncertainty regarding the familial placement of the genus

Gobiobotia paucirastella Zheng & Yan 1986    paucus, few; rastellum, diminutive of rastrum, rake, referring to fewer number of gill rakers compared to G. tungi

Gobiobotia tungi Fang 1933    in honor of L. M. Tung, West Lake Museum zoologist and professor at University of Chekiang, who loaned type specimen to Fang           

Gobiobotia yuanjiangensis Chen & Cao 1977    ensis, suffix denoting place: Yuanjiang system, Yunnan Province, China, type locality       

Subgenus Progobiobotia Chen & Cao 1977    pro-, in front of or forward, presumably referring to anterior portion of swim bladder, which is oval and unique in Gobiobotia      

Gobiobotia abbreviata Fang & Wang 1931    shortened, referring to shorter pectoral fins compared to G. pappenheimi, G. ichangensis (=filifer) and G. kiatingensis (=filifer)                

Gobiocypris Ye & Fu 1983    etymology not explained, presumably a combination of Gobio (although minnow does not resemble most other gobionines) and cypris, a small carp and a common suffix for cyprinid genera; cypris could also refer to Aphyocypris, an ex-danionine to which the authors thought this genus was related

Gobiocypris rarus Ye & Fu 1983    rare; although its scarcity is not discussed in the original description, it is quite rare

Hemibarbus Bleeker 1860    hemi-, partial, at the time believed to be allied to the genus Barbus but differing in having two barbels instead of four   

Hemibarbus brevipennus Yue 1995     brevis, short; pennus, presumably a misspelling of pinnis, fin, referring to shorter fins, especially the dorsal fin, compared to H. labeo  

Hemibarbus labeo (Pallas 1776)    one with large lips, referring to its well-developed lips

Hemibarbus longirostris (Regan 1908)    longus, long; rostris, snout, referring to long, recurved snout, 13/5 as long as eye     

Hemibarbus macracanthus Lu, Luo & Chen 1977    macro-, long; acanthus, spine, referring to long dorsal fin spine, its length 1.20-1.25 times length of head

Hemibarbus maculatus Bleeker 1871    spotted, referring to small, irregular spots on back, fins and tail             

Hemibarbus medius Yue 1995    in the middle, its main characters intermediate between H. labeo and H. maculatus               

Hemibarbus mylodon mylodon (Berg 1907)    mylo-, grinder; odon, teeth, referring to large fourth pharyngeal tooth, thickened and rounded (molariform) with a small, round excavation on upper surface          

Hemibarbus mylodon eristigma (Jordan & Hubbs 1925)    eri-, very; stigma, spot or mark, referring to “striking” coloration, comprising 7-8 large spots and smaller, blacker spots (arranged in longitudinal rows) on sides, and rows of spots on dorsal and caudal fins     

Hemibarbus qianjiangensis Yu 1990    ensis, suffix denoting place: the Qianjiang (i.e., Qian River), Zhejiang Province, China, type locality                

Hemibarbus songloensis Nguyen 2001    ensis, suffix denoting place: Sông Lô, Hà Giang Province, Viêt Nam, type locality           

Hemibarbus thacmoensis Nguyen 2001    ensis, suffix denoting place: Thac Mo, Na Hang District, Tuyên Quang Province, Viêt Nam, type locality                 

Hemibarbus umbrifer (Lin 1931)    shady, presumably referring to brownish coloration

Ladislavia Dybowski 1869    ia, belonging to: patronym not identified but almost certainly a Slavic spelling of the Polish Władysław, referring to zoologist Władysław Taczanowski (1819-1890), whose complete name is formed by the complete binomial of the only species in the genus                  

Ladislavia taczanowskii Dybowski 1869    patronym not identified but almost certainly in honor of Polish zoologist Władysław Taczanowski (1819-1890)               

Mesogobio Bănărescu & Nalbant 1973    meso-, middle, being intermediate between Gobio and other gobionine genera              

Mesogobio lachneri Bănărescu & Nalbant 1973    in honor of Ernest A. Lachner (1916-1996), curator of fishes at the U.S. National Museum, for facilitating the senior author’s visits to several museums in the United States           

Mesogobio tumenensis Chang 1980    ensis, suffix denoting place: Tumen River (boundary between China, North Korea and Russia), type locality

Microphysogobio Mori 1934    micro-, small, physinx, bladder, referring to reduced swim bladder, its anterior chamber enclosed in a thick fibrous capsule and its posterior chamber small; gobio, gudgeon

Microphysogobio alticorpus Bănărescu & Nalbant 1968    altus, high; corpus, body, referring to deeper body compared to M. brevirostris     

Microphysogobio amurensis (Taranetz 1937)    ensis, suffix denoting place: Amur River system (including Khanka Lake), Russia and China   

Microphysogobio anudarini Holcík & Pivnicka 1969    in honor of the “well known” Mongolian ichthyologist Anudarin Dashidorzhi   

Microphysogobio brevirostris (Günther 1868)    brevis, short; rostris, snout, referring to short, obtuse snout, “not much longer than the diameter of the eye”     

Microphysogobio chenhsienensis (Fang 1938)    –ensis, suffix denoting place: Chenhsien (now Shengzhou City), Zhejiang Province, China, type locality

Microphysogobio chinssuensis chinssuensis (Nichols 1926)    ensis, suffix denoting place: Chintzu, Shanxi Province, China, type locality        

Microphysogobio chinssuensis multipapillatus Bănărescu & Nalbant 1973    multi-, many; papillatus, papillose, referring to more papillae on lips compared to M. c. chinssuensis  

Microphysogobio elongatus (Yao & Yang 1977)    elongate, referring to longer, more slender body compared to A. kachekensis

Microphysogobio exilicauda (Jiang & Zhang 2013)    exilis, thin or slender; cauda, tail, referring to slender caudal peduncle

Microphysogobio fukiensis (Nichols 1926)    ensis, suffix denoting place: Fukien Province, China, type locality      

Microphysogobio hsinglungshanensis Mori 1934    ensis, suffix denoting place: Hsing-lung Shan (now Xinglong County), Jehol (now Hebei) Province, China, type locality  

Microphysogobio jeoni Kim & Yang 1999    in honor of Sang-Rin Jeon, professor at Sangmyong University (Seoul, Korea), for his contributions to the study of ichthyology in Korea       

Microphysogobio kachekensis (Oshima 1926)    ensis, suffix denoting place: Kachek River, about 40 miles above town of Kachek, Hainan Island, China, type locality  

Microphysogobio kiatingensis (Wu 1930)    ensis, suffix denoting place: Kiating (now Lo-Shan), upper Yangtze River drainage, Sichuan Province, China, type locality          

Microphysogobio koreensis Mori 1935    ensis, suffix denoting place: Korea, where it is endemic        

Microphysogobio labeoides (Nichols & Pope 1927)    oides, having the form of: “bearing a superficial resemblance” to Hemibarbus labeo       

Microphysogobio liaohensis (Qin 1987)    ensis, suffix denoting place: Liaoning Province, China, where it appears to be endemic        

Microphysogobio linghensis Xie 1986    ensis, suffix denoting place: Xiaolinghe River (sometimes shortened to Linghe), Liaoning Province, China, type locality                 

Microphysogobio longidorsalis Mori 1935    longus, long; dorsalis, of the back, referring to long dorsal fin (much longer than that of M. koreensis), which, when depressed, reaches beyond origin of anal fin     

Microphysogobio microstomus Yue 1995    micro-, small; stomus, mouth, referring to smaller mouth (and thinner lips) compared to M. linghensis

Microphysogobio nudiventris Jiang, Gao & Zhang 2012    nudus, naked; ventris, belly, referring to scaleless midventral region of body that extends slightly more than 2/3 of distance between pectoral- and pelvic-fin insertions            

Microphysogobio pseudoelongatus Zhao & Zhang 2001    pseudo-, false, i.e., although similar to M. elongatus, such an appearance is false

Microphysogobio rapidus Chae & Yang 1999    Latin rapidus, quick or swift, and Anglo-Saxon rapids, referring to shallow, fast-slowing stream habitat              

Microphysogobio tafangensis (Wang 1935)    from Tafang, Yenchow, Chekiang, China, type locality       

Microphysogobio tungtingensis (Nichols 1926)    ensis, suffix denoting place: Tungting Lake, Hunan Province, China, type locality (also occurs in Russia and Mongolia)               

Microphysogobio vietnamica Mai 1978    Vietnamese, referring to distribution in northern Viêt Nam 

Microphysogobio wulonghensis Xing, Zhao, Tang & Zhang 2011    ensis, suffix denoting place: Wulonghe River, Laiyang County, Shandong Province, China, only known area of occurrence    

Microphysogobio xianyouensis Huang, Chen & Shao 2016    ensis, suffix denoting place: Xianyou County, eastern Fujian Province, China, wherein lies type locality

Microphysogobio yaluensis (Mori 1928)    ensis, suffix denoting place: Yalu River at Tsao-ho-kou, Korea, type locality       

Microphysogobio yunnanensis (Yao & Yang 1977)    ensis, suffix denoting place: Yunnan Province (Hong River basin), China, type locality (likely also occurs in Viêt Nam part of basin)

Microphysogobio zhangi Huang, Zhao, Chen & Shao 2017    in honor of Chun-Guang Zhang, Chinese Academy of Sciences, for his “great” contributions to Chinese fish taxonomy

Paracanthobrama Bleeker 1864    para-, near, referring to Bleeker’s belief that genus belongs to a group of cyprinids he called Acanthobramae (based on the Old World leuciscine genus Acanthobrama)               

Paracanthobrama guichenoti Bleeker 1864    in honor of Antoine Alphone Guichenot (1809-1876), assistant naturalist at Musée du Jardin des Plantes a Paris          

Paraleucogobio Berg 1907    para-, near, resembling Leucogobio in general shape (differing in presence of dorsal fin spine)      

Paraleucogobio notacanthus Berg 1907    notus, back; acanthus, spine, referring to strong but flexible (at the top) dorsal fin spine        

Parasqualidus Doi 2000    para-, near, referring to close resemblance to Squalidus  

Parasqualidus maii Doi 2000    in honor of Mai Dinh Yen, Hanoi Science University, “renowned” Vietnamese freshwater fish taxonomist; type specimens were collected with his “great help” 

Placogobio Nguyen 2001    combination of placo– from Placocheilus and –gobio from Discogobio, referring to adhesive lower lip common to all three genera

Placogobio bacmeensis Nguyen & Vo 2001    ensis, suffix denoting place: Bác Mê County, Hà Giang Province, Viêt Nam, where it is endemic

Placogobio nahangensis Nguyen 2001    ensis, suffix denoting place: Na Hang District, Tuyên Quang Province, Viêt Nam, where it is endemic

Platysmacheilus Lu, Luo & Chen 1977    platys, flat or broad; cheilus, lip, referring to flat and straight lower lip             

Platysmacheilus exiguus (Lin 1932)    small, short, poor or scanty, allusion not explained, perhaps referring to elongate (i.e., slender) body or to “rudimentary” gill rakers  

Platysmacheilus longibarbatus Lu, Luo & Chen 1977    longus, long; barbatus, bearded, referring to thick barbels, which extend past edge of eye almost to operculum   

Platysmacheilus nudiventris Luo, Le & Chen 1977    nudus, naked; ventris, belly, referring to scaleless chest and abdomen   

Platysmacheilus zhenjiangensis Ni, Chen & Zhou 2005    ensis, suffix denoting place: Zhenjiang City, Jiangsu Province, China, type locality            

Pseudogobio Bleeker 1860    pseudo-, false, i.e., a false Gobio, presumably referring to previous placement of type species (P. escocinus) in Gobio and its apparent relationship with that genus                

Pseudogobio banggiangensis Nguyen 2001    ensis, suffix denoting place: Bang Giang River, Cao Bằng Province, Viêt Nam, type locality             

Pseudogobio esocinus (Temminck & Schlegel 1846)    pike-like, referring to long, large and slightly upturned snout, which resembles head of the pike (Esox)             

Pseudogobio guilinensis Yao & Yang 1977    ensis, suffix denoting place: Guilin City, Guangxi Zhuang Autonomous Region, China, type locality               

Pseudogobio vaillanti (Sauvage 1878)    patronym not identified but clearly in honor of Sauvage’s colleague, zoologist Léon Vaillant (1834-1914)        

Pseudopungtungia Mori 1935    pseudo-, false, i.e., although similar (and closely related) to Pungtungia, such an appearance is false

Pseudopungtungia nigra Mori 1935    black or dusky, probably referring to dark brown coloration (in formalin) and/or two broad blackish crossbars on dorsal, anal, ventral and caudal fins     

Pseudopungtungia tenuicorpus Jeon & Choi 1980    tenuis, slender; corpus, body, referring to characteristically slender body compared to P. nigra and Pungtungia herzi              

Pseudorasbora Bleeker 1859    pseudo-, false, i.e., although P. pusilla (=parva) resembles members of the danionine genus Rasbora, such an appearance is false

Pseudorasbora elongata Wu 1939    characterized by its elongate body          

Pseudorasbora interrupta Xiao, Lan & Chen 2007    interrupted, probably referring to incomplete lateral line      

Pseudorasbora parva parva (Temminck & Schlegel 1846)    little, referring to small size, including small head and fins    

Pseudorasbora parva uchidai Okada & Kubota 1957    in honor and celebration of the 60th birthday of Tohru Uchida, Department of Zoology, Hokkaido University, alma mater of junior author         

Pseudorasbora pugnax Kawase & Hosoya 2015    belligerent, aggressive or pugnacious, referring to how males aggressively protect their territory during spawning season (it is called “Kenka-Moroko” in Japan; kenka = fighting)

Pseudorasbora pumila Miyadi 1930    dwarf, referring to “dwarfish form of the body,” smaller than P. parva

Pungtungia Herzenstein 1892    ia, belonging to: Pungtung (Pyoktong), North Korea, type locality of P. herzi           

Pungtungia herzi Herzenstein 1892    in honor of German entomologist Alfred Otto Herz (1856-1905), who collected type       

Pungtungia hilgendorfi (Jordan & Fowler 1903)    in honor of German zoologist and paleontologist Franz Hilgendorf (1839-1904), lecturer at the Imperial Medical Academy Tokyo (1873-1876), whereupon he published articles and collected several specimens of Japanese fauna       

Pungtungia shiraii Oshima 1957    in honor of Kunihiko Shirai, Bureau of Game and Hunting of the Ministry of Agriculture and Forestry, who obtained a collection of fishes downstream of the Tame River, including type of this one, and “kindly forwarded to the writer for identification”        

Rhinogobio Bleeker 1870    rhinos, snout, i.e., a Gobio with an elongated snout          

Rhinogobio cylindricus Günther 1888    referring to “low, subcylindrical” body   

Rhinogobio hunanensis Tang 1980    ensis, suffix denoting place: Hunan Province, China, where it appears to be endemic       

Rhinogobio nasutus (Kessler 1876)    large-nosed, referring to much-produced snout    

Rhinogobio typus Bleeker 1871    serving as type of genus          

Rhinogobio ventralis Sauvage & Dabry de Thiersant 1874    ventral, referring to long ventral fins, which reach the anus  

Romanogobio Bănărescu 1961    romano, referring to “frequency and strong differentiation” (translation) of members of this genus (proposed as a subgenus of Gobio) in Romania  

Romanogobio albipinnatus (Lukasch 1933)    albus, white; pinnatus, finned, referring to white paired fins   

Romanogobio amplexilabris (Bănărescu & Nalbant 1973)    amplexus, enlacing or embracing; labrum, lip, referring to how both halves of lower lip are connected                 

Romanogobio antipai (Bănărescu 1953)    in memory of Romanian zoologist Grigore Antipa (1867-1944), who collected type           

Romanogobio banaticus (Bănărescu 1960)    icus, belonging to: Banat, historical region in Central Europe comprising parts of Romania, Serbia and Hungary bordered by the River Danube to the south, the River Tisza to the west, the River Mures¸ to the north, and the Southern Carpathian Mountains to the east         

Romanogobio belingi (Slastenenko 1934)    in honor of D. E. Beling, Director of the Dnieper Biological Station, authority on Ukranian fishes frequently cited in Slastenenko’s paper    

Romanogobio benacensis (Pollini 1816)    ensis, suffix denoting place: Lago di Benaco (Lake Garda), Verona, Italy, type locality (also occurs in Slovenia)                

Romanogobio ciscaucasicus (Berg 1932)    cis-, on this side, i.e., Ciscaucasia, referring to distribution in northern Caucasus (between the Black and Caspian Seas) region from Russia to Azerbaijan              

Romanogobio elimeius (Kattoulas, Stephanidis & Economidis 1973)    ius, pertaining to: Elimeia, ancient name of area between Grevena and Servia in Greece, type locality               

Romanogobio johntreadwelli (Bănărescu & Nalbant 1973)    in honor of John Treadwell Nichols (1883-1958), curator of fishes at the American Museum of Natural History, who first studied specimens on which this new species is based (the name Gobio nicholsi being preoccupied)             

Romanogobio kesslerii (Dybowski 1862)    in honor of German-Russian zoologist Karl Federovich Kessler (1815-1881), who identified this gudgeon as Gobio (now Romanogobio) uranoscopus in 1856            

Romanogobio macropterus (Kamensky 1901)    macro-, long; pterus, fin, probably referring to pectoral fins, which extend to or beyond base of pelvic fins           

Romanogobio parvus Naseka & Freyhof 2004    small, referring to small size (up to 66.8 mm SL) compared to other gudgeons in Gobio and Romanogobio

Romanogobio pentatrichus Naseka & Bogutskaya 1998    pente, five; trichos, hair or ray, referring to five branching rays of anal fin, a diagnostic feature of the species        

Romanogobio persus (Günther 1899)    Persia, referring to type locality in northwestern Iran (also occurs in Turkey, Armenia and Georgia) 

Romanogobio tanaiticus Naseka 2001    icus, belonging to: Tanais, ancient name for Don River, Russia and Ukraine, where this gudgeon occurs        

Romanogobio tenuicorpus (Mori 1934)    tenuis, thin; corpus, body, referring to more slender body (posteriorly compressed and less deep) compared to Gobio gobio    

Romanogobio uranoscopus (Agassiz 1828)    urano, sky; scopus, watcher, referring to obliquely placed eyes, directed upward, more towards forehead              

Romanogobio vladykovi (Fang 1943)    in honor of ichthyologist Vadim D. Vladykov (1898-1986), who identified this species as a hybrid between Gobio carpathicus and G. persa carpathorossicus (= Romanogobio kesslerii) in 1931    

Sarcocheilichthys Bleeker 1860    sarco-, flesh; cheilos, lip, referring to thick lips; ichthos, fish

Sarcocheilichthys biwaensis Hosoya 1982    ensis, suffix denoting place: Lake Biwa, Japan, where it is endemic

Sarcocheilichthys caobangensis Nguyen & Vo 2001    ensis, suffix denoting place: Cao Bàng, Cao Bàng Province, Viêt Nam, where it is endemic

Sarcocheilichthys czerskii (Berg 1914)    in honor of ornithologist A. I. Czerski (1879-1921, son of Jan Czerski, celebrated Polish geologist, naturalist and explorer), who collected type

Sarcocheilichthys davidi (Sauvage 1878)    in honor of Catholic missionary and zoologist Armand David (1826-1900), who collected type

Sarcocheilichthys hainanensis Nichols & Pope 1927    ensis, suffix denoting place: Hainan Island, China, type locality (also occurs in Laos and Viêt Nam)

Sarcocheilichthys kiangsiensis Nichols 1930    ensis, suffix denoting place: eastern Kiangsi Province, China, type locality

Sarcocheilichthys lacustris (Dybowski 1872)    lacustrine (belonging to a lake), referring to distribution in lakes in lower Amur River basin, Russia

Sarcocheilichthys nigripinnis nigripinnis (Günther 1873)    niger, black; pinnis, fin, referring to black, “or at least partly black,” fins

Sarcocheilichthys nigripinnis morii Jordan & Hubbs 1925    in honor of mammalogist Tamezo Mori, Heijo High School, Seoul, Korea, who was studying Korean vertebrates at the time and presented paratype to authors while at Stanford University

Sarcocheilichthys nigripinnis scaphignathus (Nichols 1918)    scaphi-, spade; gnathus, jaw, referring to broad, rounded and flat lower jaw

Sarcocheilichthys parvus Nichols 1930    small, a “minnowlike dwarf” just 40 mm in length

Sarcocheilichthys sinensis sinensis Bleeker 1871    ensis, suffix denoting place: Sinica (China), referring to type locality in Yangtze River

Sarcocheilichthys sinensis fukiensis Nichols 1925    ensis, suffix denoting place: Fukien Province, China, type locality

Sarcocheilichthys soldatovi (Berg 1914)    in honor of ichthyologist Vladimir Konstantinovich Soldatov (1875-1941), who collected type

Sarcocheilichthys variegatus variegatus (Temminck & Schlegel 1846)    variable, probably referring to how juvenile specimens differ in color from adults, generally lighter and “usually varied with brown on the back” (translation)

Sarcocheilichthys variegatus microoculus Mori 1927    micro-, small; oculus, eye, referring to small eyes, 5.0-5.5 in head and 2.0 in snout

Sarcocheilichthys variegatus wakiyae Mori 1927    in honor of Dr. Y. Wakia, superintendent of the Korean Government Fisheries Experiment Station, for help in the preparation of Mori’s paper

Saurogobio Bleeker 1870    sauros, lizard, probably referring to elongate, lizard-like body; gobio, gudgeon

Saurogobio dabryi dabryi Bleeker 1871    in honor of Pierre Dabry de Thiersant, fish culturist, French counsel to China, and student of Chinese fishes, who sent specimens to the Muséum d’Histoire naturelle de Paris

Saurogobio dabryi chenghaiensis Dai & Yang 2002    ensis, suffix denoting place: Chenghai Lake, Yunnan Province, China, where it is endemic

Saurogobio dumerili Bleeker 1871    in honor of August Duméril (1812-1870), herpetologist and ichthyologist, Muséum national d’Histoire naturelle (Paris), who invited Bleeker to examine Chinese cyprinids in the museum’s collection

Saurogobio gracilicaudatus Yao & Yang 1977    gracilis, slender; caudatus, tailed, referring to longer, slimmer caudal peduncle compared to congeners

Saurogobio immaculatus Koller 1927    im-, not; maculosus, spotted, referring to lack of black spots compared to spotted S. dabryi

Saurogobio lissilabris Bănărescu & Nalbant 1973    lissos, smooth; labrum, lip, referring to smooth or minutely papillose lips and mental pads compared to S. dabryi

Saurogobio xiangjiangensis Tang 1980    ensis, suffix denoting place: Xiang Jiang River system, Hunan Province, China, type locality

Squalidus Dybowski 1872    etymology not explained nor evident; possibly squalidus, rough or dirty, referring to dusky markings on silvery ground color, but more likely a combination of the cyprinid names Squaliosus and Idus (= Leuciscus), inferring that Squalidus is an intermediate between these two genera

Squalidus argentatus (Sauvage & Dabry de Thiersant 1874)    silvery, although color is described as brownish yellow (other references describe color as “pale”)

Squalidus atromaculatus (Nichols & Pope 1927)    ater, black; maculatus, spotted, probably referring to black spot at base of dorsal fin

Squalidus banarescui Chen & Chang 2007    in honor of Petru Bănărescu (1921-2009), Institute of Biology, Bucharest, for his “great” contributions to Taiwanese cyprinid taxonomy, especially the subfamily Gobioninae, between 1960 and 1973

Squalidus chankaensis chankaensis Dybowski 1872    ensis, suffix denoting place: Lake Khanka (Chanka), Amur River drainage, Russia (also occurs in China and Mongolia)

Squalidus chankaensis biwae (Jordan & Snyder 1900)    ensis, suffix denoting place: Lake Biwa, Japan, where it is endemic

Squalidus chankaensis vietnamensis Bănărescu & Nalbant 1964    ensis, suffix denoting place: Viêt Nam, referring to type locality in Phong-Thô (also occurs in Laos)

Squalidus gracilis gracilis (Temminck & Schlegel 1846)    slender, referring to slender body shape

Squalidus gracilis majimae (Jordan & Hubbs 1925)    in honor of Toyoji Majima, Imperial University of the Hokkaido

Squalidus gracilis minkiangensis Bănărescu & Nalbant 1964    ensis, suffix denoting place: Minjiang River, Fujian Province, China, type locality

Squalidus gracilis tsuchigae (Jordan & Hubbs 1925)    in honor of Yasukei Tsuchiga, a science teacher at Yamada, Japan, who collected some of the fishes examined by Jordan and Hubbs

Squalidus iijimae (Oshima 1919)    in honor of zoologist Isao lijima (also spelled Ijima, 1861-1921), Science College, Imperial University of Tokyo

Squalidus intermedius (Nichols 1929)    similar in appearance to Gnathopogon elongatus and Leucogobio (= Gnathopogon) taeniellus, “being intermediate between these two genera of convenience”

Squalidus japonicus japonicus (Sauvage 1883)    Japanese, referring to type locality at Lake Biwa, Japan

Squalidus japonicus coreanus (Berg 1906)   Korean, a subspecies of S. japonicus described from South Korea

Squalidus mantschuricus (Mori 1927)    Manchurian, referring to type locality at Fushun, China

Squalidus minor (Harada 1943)    little, probably referring to small size (32 mm)

Squalidus multimaculatus Hosoya & Jeon 1984    multi-, many; maculatus, spotted, referring to 6-12 small round dark brown spots on sides

Squalidus nitens (Günther 1873)    shining, presumably referring to silvery lower half of body and/or silvery blue band along middle of sides

Squalidus wolterstorffi wolterstorffi (Regan 1908)    in honor of German geologist, herpetologist and curator Willy Wolterstorff (1864-1943), who “received” fishes from China collected by Martin Kreyenberg, including type of this species

Squalidus wolterstorffi huapingensis Wu & Wu 1989    ensis, suffix denoting place: Huaping, Yunnan Province, China, type locality

Xenophysogobio Chen & Cao 1977    xenos, strange; physo, swim bladder, i.e., a Gobio with a strangely shaped swim bladder

Xenophysogobio boulengeri (Tchang 1929)    in honor of George A. Boulenger (1858-1937), who provided “some guidance” (translation) in the completion of Tchang’s paper

Xenophysogobio nudicorpa (Huang & Zhang 1986)    nudus, bare or naked; corpa, body, referring to scaleless body except for lateral line

Subfamily TINCINAE Tench

Tinca Garsault 1764    tautonymous with Cyprinus tinca [authorship often incorrectly given as Cuvier 1816]

Tinca tinca (Linnaeus 1758)    Latin for tench