v. 22.0 – 8 Nov. 2017  view/download PDF

Subfamily CYPRININAE (Acrossocheilus through Epalzeorhynchos)
Taxonomic note: includes taxa historically placed in subfamilies Barbinae and Labeoninae.

Aaptosyax Rainboth 1991    aaptos, invincible; syax, fish, presumably alluding to large size (up to one meter) and “especially voracious” behavior

Aaptosyax grypus Rainboth 1991    curved or hook-nosed, referring to strongly curved jaws

Acapoeta Cockerell 1910    presumably a-, not, i.e., not Capoeta (proposed as a subgenus of Capoeta)

Acapoeta tanganicae (Boulenger 1900)    of Lake Tanganyika, where it is endemic

Acrossocheilus Oshima 1919    a-, without; crosso, fringe; cheilos, lip, referring to smooth upper lip, which distinguishes it from Crossocheilus

Subgenus Acrossocheilus                                 

Acrossocheilus beijiangensis Wu & Lin 1977    ensis, suffix denoting place: Bei Jiang River, Pearl River drainage, Guandong Province, China, where it is endemic

Acrossocheilus fasciatus (Steindachner 1892)    banded, referring to five narrow dark brown transverse bands running from back to near the edge of the belly

Acrossocheilus hemispinus (Nichols 1925)    hemi-, half; spinus, spine, presumably referring to last simple ray of dorsal fin, “stiffened and serrate behind, but soft distally”

Acrossocheilus iridescens (Nichols & Pope 1927)    iridescent or rainbow-like, presumably referring to its coloration

Acrossocheilus jishouensis Zhao, Chen & Li 1997    ensis, suffix denoting place: Jishou City, Hunan Province, China, type locality

Acrossocheilus kreyenbergii (Regan 1908)    in honor of German naval officer, physician and natural history collector Martin Kreyenberg, who collected type

Acrossocheilus longipinnis (Wu 1939)    longus, long; pinnis, fin, referring to long dorsal-fin filament on some specimens

Acrossocheilus macrophthalmus Nguyen 2001    macro-, large; opthalmus, eye, referring to large eyes, as long as snout

Acrossocheilus malacopterus Zhang 2005    malacos, soft; pterus, fin, referring to spinous but slender last simple dorsal-fin ray

Acrossocheilus microstoma (Pellegrin & Chevey 1936)    multi-, many; striatus, barred, referring to high number of vertical black bars on flank compared to other barred congeners

Acrossocheilus monticola (Günther 1888)    of the mountains, referring to type locality, a mountain stream flowing into Ichang gorge of Yangtze River, China

Acrossocheilus multistriatus Lan, Chan & Zhao 2014    micro-, small; stomus, mouth, referring to very small (“très petite”) mouth

Acrossocheilus parallens (Nichols 1931)    parallel or side-by-side, presumably referring to “distinct short cross-marks, confined to the sides of the back (except in young)”

Acrossocheilus rendahli (Lin 1931)    patronym not identified but probably in honor of Hialmar Rendahl (1891-1969), zoologist and artist, who described many Chinese fishes

Acrossocheilus spinifer Yuan, Wu & Zhang 2006    spinus, spine; fero, to bear, referring to spinous and stout last simple dorsal-fin ray

Acrossocheilus stenotaeniatus Chu & Cui 1989    stenos, narrow; taeniatus, banded, referring to five crossbands on body, thinner than those on A. clivosius

Acrossocheilus wenchowensis Wang 1935    ensis, suffix denoting place: Wenchow (now Wenzhou), Zhejiang Province, China, type locality

Acrossocheilus xamensis Kottelat 2000    ensis, suffix denoting place: Nam Xan basin, northeastern Laos, type locality and only known distribution

Acrossocheilus yunnanensis (Regan 1904)    ensis, suffix denoting place: Yunnan Fu (or Yunnan-Fou, now known as Kunming), Yunnan Province, China, type locality

Subgenus Lissochilichthys Oshima 1920    lissos, smooth; cheilus, lip, presumably referring to thick, fleshy lips, but possibly also referring to the “very closely related” Lissochilus Weber & de Beaufort 1916 (a junior homonym of Lissochilus Zittel 1882 [fossil gastropods] effectively replaced by Neolissochilus); ichthys, fish

Acrossocheilus clivosius (Lin 1935)    belonging to a hill, referring to mountain stream habitat

Acrossocheilus ikedai (Harada 1943)    in honor of Kiyoshi Ikeda, General, Japanese Imperial Navy

Acrossocheilus lamus (Mai 1978)    of the Lam River, northern Viêt Nam, presumably referring to type locality and possibly only known distribution

Acrossocheilus paradoxus (Günther 1868)    strange or contrary to expectation, referring to a “very singular variation” in which one specimen had a “considerably produced” snout and three others had shorter, obtuse snouts

Acrossocheilus wuyiensis Wu & Chen 1981    ensis, suffix denoting place: Wuyishan Nature Reserve, Fujian Province, China, where it is endemic

Albulichthys Bleeker 1859    Albula, referring to how “peculiar profile” (translation) of A. albuloides reminded Bleeker of the bonefish (Albulidae) Albula bananus (=glossodonta); ichthys, fish

Albulichthys albuloides (Bleeker 1855)    oides, having the form of: referring to how its “peculiar profile” (translation) reminded Bleeker of the bonefish (Albulidae) Albula bananus (=glossodonta)

Altigena Burton 1934    altus, high; genys, chin; name coined by Lin 1934 as a subgenus of Osteochilus distinguished by its higher, deeper cheeks [Lin included multiple species without designating which one was type, so authorship dates to Burton]

Altigena binhluensis (Nguyen 2001)    ensis, suffix denoting place: Bình Lu, Phong Thô, Viêt Nam, type locality

Altigena daos (Nguyen 2001)    of the Dà River (Black River) basin, Lai Châu, Son La, Hoà Bình and part of Yên Lai Provinces, Viêt Nam, where it is endemic

Altigena discognathoides (Nichols & Pope 1927)    oides, having the form of: perhaps referring to interior transverse mouth, which authors believed was a “relic … from ancestral bottom-living fishes such as Garra” (senior synonym of, but at that time used interchangeably with, Discognathus [Heckel 1843])

Altigena elegans (Kottelat 1998)    tasteful, presumably reflecting Kottelat’s opinion about its shape, form and/or color

Altigena lippa (Fowler 1936)    bleary-eyed, presumably referring to gray iris and/or pale border around eye

Altigena loos (Nguyen 2001)    of the Lô River (also occurs in Gâm River), Hà Giang and Tuyên Quang Provinces, Viêt Nam, where it is endemic

Altigena sinkleri (Fowler 1934)    in honor of Mr. J. M. R. Sinkler of Philadelphia, USA, who helped collect fishes in Thailand

Altigena tonkinensis (Pellegrin & Chevey 1934)    ensis, suffix denoting place: Nghia Lô, Tonkin, northern Viêt Nam, type locality

Altigena wui (Zheng & Chen 1983)    in honor of Hsien-Wen Wu, National Research Institute of Biology, Academia Sinica, for his contributions to the systematics of Chinese fishes

Altigena yunnanensis (Wu & Lin 1977)    ensis, suffix denoting place: Yunnan Province, China, where it is endemic

Altigena zhui (Zheng & Chen 1989)    in honor of Y.-D. Zhu, coauthor of an unpublished 1963 manuscript in which this species was first described

Amblyrhynchichthys Bleeker 1859    amblys, blunt or obtuse, rhynchus, snout, referring to obliquely truncated snout; ichthys, fish

Amblyrhynchichthys micracanthus Ng & Kottelat 2004     micro-, small; acanthus, thorn or spine, referring to smaller dorsal spine compared to A. truncatus

Amblyrhynchichthys truncatus (Bleeker 1850)    referring to obliquely truncated snout

Arabibarbus Borkenhagen 2014    Arabi-, referring to Arabian distribution of this genus, previously included in Barbus

Arabibarbus arabicus (Trewavas 1941)    Arabian, referring to distribution in Arabian Peninsula (Yemen to Saudi Arabia)

Arabibarbus grypus (Heckel 1843)    curved, perhaps referring to how profile of brow forms a “parabolic arc down over the nose” (translation)

Arabibarbus hadhrami Borkenhagen 2014     derived from its native range, the Hadhramaut Province in Yemen, in honor of the people who live there and call themselves “Hadhrami”

Aspiorhynchus Kessler 1879    aspio-, referring to leuciscine genus Aspius Agassiz 1832, which also has a wedge-shaped head and rhynchus, snout

Aspiorhynchus laticeps (Day 1877)    latus, wide; ceps, head, referring to broad upper surface of head, its width nearly twice its height

Aulopyge Heckel 1841    aulos, flute; pyge, behind or rump, i.e., “fluted tail stem,” referring to prolonged urogenital papilla fused with anterior edge of anal fin on female

Aulopyge huegelii Heckel 1843    patronym not identified but probably in honor of Austrian naturalist and diplomat Charles von Hügel (1795-1870), who collected many fishes for Heckel, usually in India (but presumably not this one, which occurs in Bosnia and Herzegovina and Croatia)

Balantiocheilos Bleeker 1859    balantion, pouch; cheilus, lip, referring to posterior margin of lower lip, which forms a pouch or pocket that opens posteriorly between lip and skin of throat

Balantiocheilos ambusticauda Ng & Kottelat 2007    ambustus, burned around or scorched; cauda, tail, referring to black edge of caudal fin

Balantiocheilos melanopterus (Bleeker 1850)    melanos, black; pterus, fin, referring to black along distal margins of dorsal, caudal, anal, and pelvic fins

Bangana Hamilton 1822    Banggana, native Bengali name “common to most species” of Mugil (Mugilidae) and to certain cyprinids, which share an “elevated longitudinal ridge on the middle of the lower jaw”

Bangana almorae (Chaudhuri 1912)    of Almora, western Himalayas, India, type locality

Bangana ariza (Hamilton 1807)    presumably derived from Arija, Telugu (official language of Andhra Pradesh, India) name for this fish

Bangana brevirostris Liu & Zhou 2009    brevis, short; rostrum, snout, referring to “relatively short snout”

Bangana decora (Peters 1881)    beautiful, allusion not evident, perhaps referring to golden scales edged with black

Bangana dero (Hamilton 1822)    presumably local Bengali name for this fish (also spelled doro)

Bangana devdevi (Hora 1936)    in honor of Dev Dev Mukerji, Technical Assistant, Zoological Survey of India, who first noted this species was distinct from B. dero

Bangana diplostomus (Heckel 1838)    diplo, double; stomus, mouth, presumably referring to how “lower lip lays flat behind the edge of the lower jaw, so at first sight, its finely serrated bow seems to be the real mouth opening” (translation) when mouth is closed

Bangana gedrosicus (Zugmayer 1912)    icus, belonging to: Gedrosia, ancient name of Balochistan Province, Pakistan, including Panjgur, type locality

Bangana lemassoni (Pellegrin & Chevey 1936)    in honor of Jean L. Lemasson, Engineer of Water and Forests (Hanoi, Viêt Nam), who collected type

Bangana rendahli (Kimura 1934)    in honor of zoologist and artist Hialmar Rendahl (1891-1969), who had identified this cyprinid in 1933 as Labeo diplostomus (=B. diplostoma)

Bangana tungting (Nichols 1925)    referring to Tungting Lake, Hunan Province, China, type locality

Bangana xanthogenys xanthogenys (Pellegrin & Chevey 1936)    xanthos, yellow; genys, chin, referring to cheek coloration

Bangana xanthogenys songloensis (Nguyen 2001)    ensis, suffix denoting place: Sông Lô, Tuyên Quang Province, Viêt Nam, type locality

Barbichthys Bleeker 1859    referring to previous placement of B. laevis in Barbus; ichthys, fish

Barbichthys laevis (Valenciennes 1842)    smooth, perhaps referring to “radius of the dorsal [which] is smooth and without perforation” (translation)

Barbodes Bleeker 1859    iodes, having the form of: presumably referring to similarity to and/or affinity with Barbus

Barbodes amarus (Herre 1924)    bitter, from the Moro (Philippines) pait, a word used to describe the bitter taste of smaller East Indian cyprinids

Barbodes aurotaeniatus (Tirant 1885)    auro, gold; taeniatus, banded, referring to brilliant golden-yellow band atop the lateral line

Barbodes banksi Herre 1940    in honor of Edward Banks (1903-1988), curator, Sarawak Museum, “without whose kindly aid little could have been done during [Herre’s] brief stay in Kuching”

Barbodes baoulan Herre 1926    Bao-u-lan, Marinão name for this species in the Philippines

Barbodes binotatus (Valenciennes 1842)    bi-, two; notatus, marked, referring to large blotch at anterior base of dorsal fin and round spot in middle of caudal peduncle

Barbodes bovanicus (Day 1877)    icus, belonging to: Bowany River (Madras, India), type locality (spelling is latinized, with v replacing w)

Barbodes bunau (Rachmatika 2005)    vernacular for this species in the language of Dayak Punan, one of the native peoples living in the Seturan watershed, East Kalimantan, Borneo (type locality)

Barbodes cataractae (Fowler 1934)    of the cataract or cascade, referring to Cascade River, Murcielagos Bay, Mindanao, Philippines, type locality

Barbodes clemensi Herre 1924    in honor of Herre’s friend Chaplain Joseph Clemens, who with his wife made the first scientific collections around Lake Lanao, Mindanao, Philippines, only known area of occurrence

Barbodes disa Herre 1932    Marinão name for this species in the Philippines

Barbodes dunckeri (Ahl 1929)    in honor of ichthyologist Georg Duncker (1870-1953), Zoological Museum Hamburg, who recognized the distinctiveness of, but did not name, this cyprinid in 1905

Barbodes elongatus (Oshima 1920)    referring to its elongate body

Barbodes everetti (Boulenger 1894)    patronym not identified but clearly in honor of Alfred Hart Everett (1848-1898), British civil servant and administrator in Borneo and naturalist who collected type

Barbodes flavifuscus (Herre 1924)    flavus, yellow; fuscus, dusky, probably referring to yellowish brown belly coloration in alcohol and perhaps also dusky fins

Barbodes hemictenus Jordan & Richardson 1908    hemi-, half; ktenos, comb, allusion not evident, perhaps referring to weaker and fewer serratures on third dorsal spine compared to Barbodes maculatus (=binotatus)

Barbodes herrei (Fowler 1934)    in honor of ichthyologist-lichenologist Albert W. Herre (1868-1962), for his many contributions to Philippine ichthyology

Barbodes ivis (Seale 1910)    presumably local name for this species at Balabac Island, Philippines, type locality

Barbodes joaquinae (Wood 1968)    in honor of Joaquina C. Wood, “who first collected this fish during her many trips afield while studying Mindanao fauna”

Barbodes katolo Herre 1924    Moros name for this species in the Philippines

Barbodes kuchingensis (Herre 1940)    –ensis, suffix denoting place: 18 miles east of Kuching, Sarawak state, Borneo, type locality

Barbodes lanaoensis Herre 1924    –ensis, suffix denoting place: Lake Lanao, Mindanao, Philippines, type locality

Barbodes lateristriga (Valenciennes 1842)    lateris; side; striga, streak, presumably referring to two short vertical bars on sides

Barbodes lindog Herre 1924    Moros name for this species in the Philippines

Barbodes manguaoensis (Day 1914)    –ensis, suffix denoting place: Lake Manguao, northern Palawan Island, Philippines, type locality

Barbodes manalak Herre 1924    Moros (Philippines) vernacular applied to the two largest cyprinid species of Lake Lanao, including this one

Barbodes microps (Günther 1868)    micro-, small; ops, eye, referring to much smaller eye compared to Barbus maculatus (=Barbodes binotatus)

Barbodes montanoi (Sauvage 1881)    in honor of French ethnologist Joseph Montano (1844-?), who collected or secured type

Barbodes pachycheilus (Herre 1924)    pachys, thick; cheilus, lip, referring to broad, thick, fleshy upper lip

Barbodes palaemophagus (Herre 1924)    Palaemon, a sea god and a genus of caridean shrimps; phagus, eating, referring to a Palaemon in mouth and throat of type specimen, “which it had evidently been in the act of swallowing when captured”

Barbodes palata Herre 1924    Marinao word for narrow, applied to this fish by the Moros because of its pinched narrow head, especially along ventral side           

Barbodes palavanensis (Boulenger 1895)    –ensis, suffix denoting place: Palawan Island, Philippines, type locality

Barbodes polylepis Chen & Li 1988    poly, many; lepis, scale, referring to 45-46 scales along lateral line

Barbodes quinquemaculatus (Seale & Bean 1907)    quinque, five; maculatus, spotted, referring to distinct round black spot on base of caudal, another at origin of dorsal, a third less distinct at origin of anal, and two round black spots on median line near middle of body

Barbodes resimus (Herre 1924)    turned upward, referring to “back very strongly arched from head to origin of dorsal” [also spelled resinus by Herre, presumably in error]

Barbodes rhombeus (Kottelat 2000)    rhombic, allusion not explained, perhaps referring to shape of spots and blotches on body and caudal peduncle

Barbodes sealei Herre 1933    in honor of ichthyologist Alvin Seale (1871-1958), Stanford University, who described this barb in 1910 but used a preoccupied name

Barbodes semifasciolatus (Günther 1868)    semi-, half; fasciolatus, banded referring to 3-4 narrow, black vertical streaks on sides and tail

Barbodes sirang Herre 1932    Marinão name for this species in the Philippines

Barbodes stigma (Valenciennes 1844)    spot or mark, referring to small black spot on fourth and fifth soft rays of dorsal fin    

Barbodes tras Herre 1926    Marinão name for this species in the Philippines

Barbodes truncatulus (Herre 1926)    truncated, referring to “truncate form of lower jaw”

Barbodes tumba Herre 1924    Marinao or Lanao Moro (Philippines) name for several species of cyprinids, including, apparently, this one

Barbodes umalii (Wood 1968)    in honor of Agustin F. Umali, Senior Ichthyologist, National Museum of the Philippines, who encouraged Wood to study fishes in Mindanao, for “his vast knowledge of prewar ichthyology and long hours spent passing this information on to the writer”

Barbodes wynaadensis (Day 1873)    ensis, suffix denoting place: Wynaad, Kerala, India, type locality

Barbodes xouthos (Kottelat & Tan 2011)    golden yellow, referring to yellow-brown coloration in life and/or yellow pelvic fin in adults

Barboides Brüning 1929    oides, having the form of: Barbus, i.e., resembling a barb

Barboides britzi Conway & Moritz 2006    in honor of Ralph Britz (Natural History Museum, London), whose “contagious enthusiasm for ichthyology and interest in tiny fishes has left a lasting impression” on both authors

Barboides gracilis Brüning 1929    slender or thin, referring to its “delicate shape” (translation)

Barbonymus Kottelat 1999    Barbus, generic name earlier applied to these fishes; anonymus, without name, referring to a group of southeast Asian species that had gone without a proper generic name until now

Barbonymus altus (Günther 1868)    high, presumably referring to its elevated body

Barbonymus balleroides (Valenciennes 1842)    oides, having the form of: “small bream” (translation), presumably the Old World leuciscin Ballerus ballerus

Barbonymus belinka (Bleeker 1860)    local vernacular for this species in Sumatra, Indonesia

Barbonymus collingwoodii (Günther 1868)    in honor of surgeon-naturalist Cuthbert Collingwood (1826-1908), who presented type to the British Museum (Natural History)

Barbonymus gonionotus (Bleeker 1849)    gonio-, angle; notos, back, referring to arched dorsal profile

Barbonymus mahakkamensis (Ahl 1922)    –ensis, suffix denoting place: Mahakam River, eastern Kalimantan Province, eastern Borneo, type locality

Barbonymus platysoma (Bleeker 1855)    platy, flat; soma, body, referring to strongly compressed body

Barbonymus schwanenfeldii (Bleeker 1854)    in honor of military surgeon H. W. Schwanefeld, who collected type (Bleeker misspelled the name, a mistake he subsequently corrected, but original spelling must be retained)

Barbonymus strigatus (Boulenger 1894)    furrowed or grooved, probably referring to seven dark stripes along body, which appear to form a furrow between rows of scales

Barbonymus sunieri (Weber & de Beaufort 1916)    in honor of Armand Louis Jean Sunier (1886-1974), Dutch biologist and later director of Artis Zoo Amsterdam

Barbopsis Di Caporiacco 1926    opsis, likeness, referring to similarity to Barbus

Barbopsis devecchii Di Caporiacco 1926    in honor of Cesare Maria De Vecchi (1884-1959), governor of Italian Somaliland (now a region of Somalia), where it is endemic

Barbus Cuvier & Cloquet 1816    barbel, tautonymous with Cyprinus barbus: referring to four barbels, two at tip at one at each corner of mouth, and perhaps also to its vernacular (e.g., barbylle in Middle English, barbel in Old French)

Barbus balcanicus Kotlík, Tsigenopoulos, Ráb & Berrebi 2002    icus, belonging to: Balkan Peninsula, where it is widely, and almost exclusively, distributed

Barbus barbus (Linnaeus 1758)    barbel, referring to four barbels, two at tip at one at each corner of mouth, and perhaps also to its vernacular (e.g., barbylle in Middle English, barbel in Old French)

Barbus bergi Chichkoff 1935    patronym not identified but undoubtedly in honor of Soviet ichthyologist Lev (or Leo) Semyonovich Berg (1876-1950)

Barbus biharicus Antal, László & Kotlík 2016    icus, belonging to: Bihar Counties (in both Romania and Hungary), where it is endemic

Barbus borysthenicus Dybowski 1862    icus, belonging to: Borysthenes, ancient name for Dnieper River, referring to type locality in the Ukraine

Barbus caninus Bonaparte 1839    dog, from Barb canin, local name for this barb in Piedmont region of Italy

Barbus carpathicus Kotlík, Tsigenopoulos, Ráb & Berrebi 2002    icus, belonging to: Carpathian Mountains, the northwestern part of which the distribution of this species is restricted

Barbus ciscaucasicus Kessler 1877    cis-, on this side, referring to distribution in northern Caucasus (between the Black and Caspian Seas) region of Russia (Dagestan, Azerbaijan)

Barbus cyclolepis Heckel 1837    cyclo-, circle; lepis, scale, referring to smaller, rounder scales compared to B. communis (=barbus)

Barbus cyri De Filippi 1865    of the Kura River (Cyrus in Latin), Georgia, type locality

Barbus ercisianus Karaman 1971    anus, belonging to: Erciş, Van Province, Turkey, type locality

Barbus euboicus Stephanidis 1950    icus, belonging to: Evia (=Eúboia) Island, Greece, where it is endemic

Barbus goktschaicus Kessler 1877    icus, belonging to: Sevan (=Goktscha) Lake, Armenia, type locality

Barbus haasi Mertens 1925    in honor of malacologist Fritz Haas (1886-1969), who collected type in the Mediterranean waters of Spain

Barbus karunensis Khaefi, Esmaeili, Geiger & Eagderi 2017   ensis, suffix denoting place: Karun River, Iran, type locality

Barbus kubanicus Berg 1912    icus, belonging to: Kuban River, Russia, type locality

Barbus lacerta Heckel 1843    lizard, allusion not explained nor evident

Barbus macedonicus Karaman 1928    Macedonian, referring to type locality in Vardar River, Macedonia (also occurs in Greece)

Barbus meridionalis Risso 1827    southern, presumably referring to distribution in southern Europe and/or southern France

Barbus miliaris De Filipi 1863    arius, pertaining to: mille, thousand, presumably referring to fine black speckles (i.e., appearing like a thousand dots) on belly and dorsal and anal fins

Barbus niluferensis Turan, Kottelat & Ekmekç 2009    ensis, suffix denoting place: Nilüfer River drainage, Turkey, type locality

Barbus oligolepis Battalgil 1941    oligo-, few; lepis, scale, presumably referring to fewer number of scales compared to B. tauricus polylepis (=cyclolepis) described in same publication

Barbus peloponnesius Valenciennes 1842    Peloponnesian, referring to type locality in Morée (=Pelopponese), Greece

Barbus petenyi Heckel 1852    in honor of Hungarian ornithologist Johann Petényi, 1799-1855 (barb is from nearby Romania and Bulgaria)

Barbus plebejus Bonaparte 1839    plebius, common, allusion not explained; name first appeared in Valenciennes (1829) and may refer its being a variant of B. vulgaris (=barbus), whose name also means common

Barbus prespensis Karaman 1924    ensis, suffix denoting place: Lake Prespa and tributaries, Republic of Macedonia, where it is endemic

Barbus rebeli Koller 1926    patronym not identified but probably in honor of lepidopterist Hans Rebel (1861-1940), who became general director of the Naturhistorisches Museum in Vienna in 1925

Barbus sperchiensis Stephanidis 1950    ensis, suffix denoting place: Sperchios (Spercios) River, Greece, type locality

Barbus strumicae Karaman 1955    of the Strumica, a river in Macedonia, type locality (also occurs in Greece and Bulgaria)

Barbus tauricus Kessler 1877    icus, belonging to: Taurica, ancient Greek and Roman name for Crimea, being the only Barbus in the Crimean Peninsula

Barbus thessalus Stephanidis 1971     of Thessaly, Pinios River, Greece, where it is endemic

Barbus tyberinus Bonaparte 1839    inus, belonging to: Tiber (Tevere) River, type locality near Rome, Italy

Barbus waleckii Rolik 1970   in honor of Antoni Wałecki (1815-1897), Polish zoologist who noted enormously large specimens of B. cyclolepis (misidentified as B. petenyi) in the Vistula River near Warsaw in 1864, now recognized as this species

Caecobarbus Boulenger 1921    caecus, blind, i.e., a blind (eyeless) Barbus

Caecobarbus geertsii Boulenger 1921    in honor of amateur cave explorer G. Geerts, railway director, who collected type specimens and donated them to the Musée du Congo belge

Caecocypris Banister & Bunni 1980    caecus, blind, referring to its vestigial and sightless eyes; cypris, a small carp, a common suffix for cyprinid genera

Caecocypris basimi Banister & Bunni 1980    in honor of Basim Al-Azzawi, Natural History Research Centre, University of Baghdad, who collected type

Capoeta Valenciennes 1842    tautonymous with Cyprinus capoeta (see C. capoeta for etymology)

Capoeta aculeata (Valenciennes 1844)    sharp-pointed or stinging, possibly referring to serrated dorsal fin ray

Capoeta alborzensis Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio 2016    –ensis, suffix denoting place: Alborz Mountains (Tehran Province, Iran), where Nam River (type locality) originates

Capoeta anamisensis Zareian, Esmaeili & Freyhof 2016    –ensis, suffix denoting place: Anamis, old name of Minab city and the Minab River, Iran, where it occurs

Capoeta angorae (Hankó 1925)    of Angora, historic name of Ankara, Turkey, referring to distribution in Turkey

Capoeta antalyensis (Battalgil 1943)    ensis, suffix denoting place: Antalya, southwestern Turkey, type locality

Capoeta aydinensis Turan, Küçük, Kaya, Güçlü & Bektaş 2017    ensis, suffix denoting place: Aydin, city and eponymous province in Turkey, type locality

Capoeta baliki Turan, Kottelat, Ekmekçi & Imamoglu 2006    in honor of Süleyman Balik, for his contributions to the knowledge of Turkish fishes

Capoeta banarescui Turan, Kottelat, Ekmekçi & Imamoglu 2006    in honor of Petru Bănărescu (1921-2009), Institute of Biology, Bucharest, for his contributions to the knowledge of Turkish fishes

Capoeta barroisi Lortet 1894    in honor of Théodore Barrois, La Faculté de Médecine de Lille, who collected type and whose monograph on Syrian lakes contained this description

Capoeta bergamae Karaman 1969    of Bergama, İzmir Province, western Turkey, type locality

Capoeta buhsei Kessler 1877    in honor of botanist Friedrich Alexander Buhse (1821-1898), who collected type

Capoeta caelestis Schöter, Özuluğ& Freyhof 2009    heavenly, named after Göksu River, type locality (Turkish: gök, heavenly; su, water)

Capoeta capoeta (Güldenstädt 1773)    derived from kapwaeti, Armenian and Georgian name for female C. capoeta packed with eggs

Capoeta coadi Alwan, Zareian & Esmaeili 2016    in honor of Brian W. Coad (Canadian Museum of Nature, Ottawa), for his valuable contribution to the knowledge of the freshwater fishes of Iran

Capoeta damascina (Valenciennes 1842)    ina, belonging to: Damascus, Syria, type locality

Capoeta ekmekciae Turan, Kottelat, Kirankaya & Engin 2006    in honor of F. Güler Ekmekçi, for her contribution (e.g., help with fieldwork) to the authors’ research on the fishes of Anatolia

Capoeta erhani Turan, Kottelat & Ekmekçi 2008    in honor of Erhan Ünlü, for his contribution to the authors’ research on the fishes of Anatolia

Capoeta ferdowsii Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017    in honor of Abu Al-Qasim Ferdowsi Tusi (935-1020), Persian poet and the author of Shahnameh, the world’s longest epic poem created by a single poet, and the national epic of Greater Iran

Capoeta fusca Nikolskii 1897    dark or dusky, referring to dark coloration on back and upper sides

Capoeta kosswigi Karaman 1969    in honor of Turkish zoologist and geneticist Curt Kosswig (1903-1982), who collected type

Capoeta mandica Bianco & Bănărescu 1982    ica, belonging to: Mand River, near Dasht-e-Arzhan, Persian Gulf basin, Iran, type locality

Capoeta mauricii Küçük, Turan, Şahin & Gülle 2009    in honor of Swiss ichthyologist Maurice Kottelat (b. 1957), for his contribution to the knowledge of European and Asian fishes

Capoeta pestai (Pietschmann 1933)    patronym not identified but probably in honor of crustacean biologist Otto Pesta (1885-1974), Pietschmann’s colleague at the Natural History Museum of Vienna

Capoeta pyragyi Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017    in honor of Magtymguly Pyragy (1724-1783), Turkmen spiritual leader and philosophical poet

Capoeta razii Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio 2017    in honor of Abū Bakr Muhammad ibn Zakariyyā al-Rāzī (845-925), Persian polymath, physician, alchemist, and philosopher, for his “important” contributions in the history of medicine

Capoeta saadii (Heckel 1847)    of Saadi, a spring, presumably at or near Persepolis, Pulwar River, Kor River basin, ruins northeast of Shiraz, Iran, type locality

Capoeta sevangi De Filippi 1865    of Lake Sevanga (Russian name of Lake Sevan), Armenia, where it is endemic

Capoeta shajariani Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017    in honor of Mohammad-Reza Shajarian (b. 1940), “acclaimed” Iranian classical singer, composer and master of Persian traditional music

Capoeta sieboldii (Steindachner 1864)    patronym not identified but probably in honor of physiologist and zoologist Karl (or Carl) Theodor Ernst von Siebold (1804-1885), who expanded the fish collection at the Bavarian State Collection of Zoology (München, Germany)

Capoeta tinca (Heckel 1843)    etymology not explained nor evident, probably referring to resemblance of some physical attribute(s) to that of the Tench, Tinca tinca

Capoeta trutta (Heckel 1843)    referring to resemblance of its small scales and x-shaped spots to those of the Brown Trout, Salmo trutta

Capoeta turani Özuluğ & Freyhof 2008    in honor of Davut Turan, author of “important” papers on Anatolian Capoeta

Capoeta umbla (Heckel 1843)    referring to resemblance of its small scales and “tender beautiful nature” (translation) to that of the char Salmo (=Salvelinus) umbla

Capoetobrama Berg 1916    etymology not explained; apparently a combination of Capoeta (derived from kapwaeti, Armenian and Georgian name for female Capoeta capoeta packed with eggs), perhaps referring to reports of heavily gravid female Capoetobrama kuschakewitschi, and Acanthobrama (Leuciscinae), to which Berg believed this genus was closely related

Capoetobrama kuschakewitschi kuschakewitschi (Kessler 1872)    in honor A. A. Kushakewitsch, traveler and explorer through Middle Asia, who collected type

Capoetobrama kuschakewitschi orientalis Nikolskii 1934    eastern, referring to distribution in Chu River, Kazakstan, in the eastern end of the Aral Sea basin

Carasobarbus Karaman 1971    etymology not explained, presumably caraso-, referring to similarity to carps (Carassius) and previous placement of C. luteus in Barbus (i.e., carp-like barbs)

Carasobarbus apoensis (Banister & Clarke 1977)    ensis, suffix denoting place: a-, without; poa, grass, “coming from a grassless place,” referring to Arabian desert habitat

Carasobarbus canis (Valenciennes 1842)    dog, Latin translation of Kaelb, vernacular for this species in the Jordan River area of what is now Palestine

Carasobarbus chantrei (Sauvage 1882)    in honor of anthropologist and archaeologist Ernest Chantre (1843-1924), who collected type

Carasobarbus exulatus (Banister & Clarke 1977)    banished, referring to isolated habitats in Wadi (=valley) Hadramut and Wadi Maran, Yemen

Carasobarbus kosswigi (Ladiges 1960)    in honor of Turkish zoologist and geneticist Curt Kosswig (1903-1982), who collected type in 1939 and informed Ladiges of its uniqueness

Carasobarbus luteus (Heckel 1843)    yellow, referring to main coloration of “soft yellow” (translation)

Carasobarbus moulouyensis (Pellegrin 1924)    ensis, suffix denoting place: Moulouya basin, Morocco, type locality

Carasobarbus sublimus (Coad & Najafpour 1997)    exalted, named for type locality, A’la River, Khuzestan Province, southern Iran (A’la=most high or exalted)

Carassioides Oshima 1926    oides, having the form of: Carassius, reflecting Oshima’s belief that it is the “nearest relative” of Carassius

Carassioides acuminatus (Richardson 1846)    sharpened or pointed, presumably referring to its “elevated back, shaped in profile like the roof of a house, with the summit at the commencement of the dorsal”

Carassius Jarocki 1822    tautonymous with Cyprinus carassius, from the French carassin, carp

Carassius auratus auratus (Linnaeus 1758)    gilded, referring to golden color

Carassius auratus argenteaphthalmus Nguyen 2001    argentea, silvery; opthalmus, eye, referring to white (not red) ring around eyes

Carassius auratus burgeri Temminck & Schlegel 1846    patronym not identified but clearly in honor of physicist and biologist Heinrich Bürger (ca. 1806-1858), who collected many of the Japanese specimens described by Temminck and Schlegel

Carassius auratus grandoculis Temminck & Schlegel 1846    grand, large; oculus, eye, referring to larger eyes compared to C. a. burgeri and C. cuvieri

Carassius carassius (Linnaeus 1758)    from the French carassin, carp

Carassius cuvieri Temminck & Schlegel 1846    patronym not identified but clearly in honor of Georges Cuvier (1769-1832), whose work on fishes culminated with his contributions to Histoire naturelle des poissons (1828-1831)

Carassius gibelio (Bloch 1782)    from the German name for Crucian Carp, giebel

Carassius langsdorfii Temminck & Schlegel 1846    patronym not identified but probably in honor of Georg Heinrich von Langsdorff (1774-1852), Prussian naturalist and diplomat in Japan, who collected and/or supplied type

Carassius praecipuus Kottelat 2017    Latin for “which is not common,” referring to unexpected presence of a Carassius in the Mekong drainage and its very low dorsal-fin ray, lateral-line scale and gill-raker counts

Catlocarpio Boulenger 1898    catlo, reflecting Boulenger’s belief that it is “allied” to Catla Valenciennes 1844 (=Gibelion Heckel 1843) based on similarity in structure of mouth; carpio, carp

Catlocarpio siamensis Boulenger 1898    ensis, suffix denoting place: Siam, referring to type locality in Menam (Chao Phya) River, Thailand

Chagunius Smith 1938    latinization of Chaguni, local Bengali name for C. chagunio in India

Chagunius baileyi Rainboth 1986    in honor of ichthyologist Reeve M. Bailey (1911-2011), University of Michigan, co-chair of Rainboth’s Ph.D. committee

Chagunius chagunio (Hamilton 1822)    from Chaguni, local Bengali name for this fish in India

Chagunius nicholsi (Myers 1924)    in honor of John Treadwell Nichols (1883-1958), curator of fishes at the American Museum of Natural History, in “slight appreciation of his generous help and interest” in Myers’ work

Chuanchia Herzenstein 1891    ia, belonging to: Chuanche (or Chuan Che, “Yellow”) River, Qinuhai Province, China, type locality

Chuanchia labiosa Herzenstein 1891    large-lipped, referring to “rather fleshy lips” (translation)

Cirrhinus Oken 1817    latinization of Cuvier’s French name for genus, Les Cirrhines, from cirrhus, fringe, referring to fringed upper lip of C. cirrhosus

Cirrhinus cirrhosus (Bloch 1795)    full of curls, referring to fringed upper lip

Cirrhinus fulungee (Sykes 1839)    Marathi vernacular for this species in India, which, as Sykes explained in 1841, he adopted “so that naturalists who travel the country can always obtain” the species

Cirrhinus jullieni Sauvage 1878    in honor of J. Jullien (no other information available), who collected type

Cirrhinus microlepis Sauvage 1878     micro-, small; lepis, scale, referring to small scales, 53-60 along lateral line

Cirrhinus molitorella (Valenciennes 1844)    diminutive of molitris, presumably reflecting Valenciennes’ belief that this species is a smaller relative of Leuciscus (=Hypophthalmichthys) molitrix

Cirrhinus mrigala (Hamilton 1822)    local Sanskrit name for this species

Cirrhinus prosemion (Fowler 1934)    pro-, before; semeion, banner, referring to “advanced” dorsal fin (a character not mentioned in description nor evident in illustration)

Cirrhinus reba (Hamilton 1822)    presumably local Bengali name for this species

Cirrhinus rubirostris Roberts 1997    ruber, red; rostrum, snout, referring to color of rostral tubercles and snout; also refers to Karen (Thai dialect) name, niya gwoh nadee, “red-nosed fish”

Clypeobarbus Fowler 1936    a Barbus with clypeus, shield, presumably referring to “very deeply exposed” scales on lateral line

Clypeobarbus bellcrossi (Jubb 1965)    in honor of Graham Bell-Cross, Zambia Department of Game and Fisheries, for his systematic study of fishes from headwater tributaries of the Zambezi River

Clypeobarbus bomokandi (Myers 1924)    referring to Bomokandi River (Democratic Republic of the Congo), type locality

Clypeobarbus breviclipeus Stiassny & Sakharova 2016    brevis, small or brief; clypeus, shield, referring to relatively small size of pore-bearing scales in midlateral series as compared with greatly enlarged, shield-like scales of some congeners

Clypeobarbus congicus (Boulenger 1899)    icus, belonging to: Congo River basin of Zaire (now Democratic Republic of the Congo), type locality

Clypeobarbus hypsolepis (Daget 1959)    hypso-, high; lepis, scale, referring to especially tall (or deep) lateral line scales, particularly below dorsal fin

Clypeobarbus matthesi (Poll & Gosse 1963)    in honor of Hubert Matthes, Royal Museum of Central Africa, who collected type

Clypeobarbus pleuropholis (Boulenger 1899)    pleuro-, side; pholis, spot or fleck, allusion uncertain, possibly referring to scales edged with crimson

Clypeobarbus pseudognathodon (Boulenger 1915)    pseudo-, false; gnathus, jaw; odon, tooth, referring to tubercles that line mouth, which appear like false teeth

Clypeobarbus schoutedeni (Poll & Lambert 1961)    in honor of Henri Schouteden, honorary director of the Musée Royal de l’Afrique centrale, on the occasion of his 80th birthday, for his tireless zoological work, and for his valuable assistance as director of Revue de Zoologie et de Botanique africaines

Cophecheilus Zhu, Zhang, Zhang & Han 2011    kopheia, depression, referring to shallow, arched, subdistal depression along ventral margin of rostral cap; cheilos, lip, a commonly used suffix for labeonine genera

Cophecheilus bamen Zhu, Zhang, Zhang & Han 2011    Ba Men, local name of this and similar species in Jingxi County, Guangxi Province, South China

Cophecheilus brevibarbatus He, Huang, He & Yang 2015    brevis, short; barbatus, bearded, referring to shorter barbels compared to C. bamen

Coptostomabarbus David & Poll 1937    a Barbus with copto-, cut or chopped small, stoma, mouth, presumably referring to upturned mouth, distinguishing feature of genus

Coptostomabarbus bellcrossi Poll 1969    in honor of Graham Bell-Cross, Zambia Department of Game and Fisheries, who helped collect type

Coptostomabarbus wittei David & Poll 1937    in honor of herpetologist Gaston François De Witte (1897-1980), who collected type

Cosmochilus Sauvage 1878    cosmos, ornament; cheilus, lip, referring to thick, fringed lips, entirely covered by large papillae

Cosmochilus cardinalis Chu & Roberts 1985    red, referring to bright red fins

Cosmochilus falcifer Regan 1906    falcatus, sickle-shaped; fero, to bear, i.e., sickle-bearing, probably referring to enlarged and serrate dorsal-fin ray

Cosmochilus harmandi Sauvage 1878    in honor of François-Jules Harmand (1845-1921), physician, explorer and diplomat, who collected type

Cosmochilus nanlaensis Chen, He & He 1992    ensis, suffix denoting place: Nanla River, Mekong River system, Yunnan Province, China, type locality

Crossocheilus Kuhl & van Hasselt 1823    crosso, fringe; cheilos, lip, referring to fringed upper lip of C. oblongus

Crossocheilus atrilimes Kottelat 2000    ater, black; limes, a narrow or elongate space, referring to midlateral stripe extending to posterior extremity of median caudal-fin rays

Crossocheilus cobitis (Bleeker 1854)    from the Greek kobitis, “like a gudgeon,” referring to loach (Cobitis)-like shape

Crossocheilus elegans Kottelat & Tan 2011    elegant, presumably referring to distinctive color pattern consisting of a blackish midlateral stripe extending from tip of gill opening to middle of base of caudal fin, separate from dark brown dorsum by a pale yellowish brown stripe

Crossocheilus gnathopogon Weber & de Beaufort 1916    gnathos, jaw; pogon, beard, presumably referring to pair of maxillary barbels

Crossocheilus langei Bleeker 1860    in honor of E. A. Lange, acting health officer and hospital inspector, Dutch East Indian Army, who forwarded type to Bleeker

Crossocheilus microstoma Ciccotto & Page 2017    micro-, small; stoma, mouth, referring to its narrower mouth compared to congeners

Crossocheilus nigriloba Popta 1904    niger, black; lobus, lobe, referring to large black spot on lower lobe of caudal fin

Crossocheilus oblongus Kuhl & van Hasselt 1823    oblong, referring to elongate, fusiform body

Crossocheilus obscurus Tan & Kottelat 2009     dark, referring to dark general appearance compared to southeast Asian congeners

Crossocheilus reticulatus (Fowler 1934)    netted, referring to reticulated pattern formed by scales with dark or blackish-brown edges

Crossocheilus tchangi Fowler 1935    in honor of Tchunlin (or Tchung-Lin) Tchang (1897-1963), Beijing Normal University, for his work on Chinese cyprinoids

Cyclocheilichthys Bleeker 1859    unneeded replacement name (now a nomen protectum) for Cyclocheilos Bleeker 1859, which Bleeker may have thought was preoccupied; cyclo-, round or circular, cheilos, lips, allusion not specified, possibly referring to continuous lips forming round, sucker-like shape when open; ichthys, fish

Cyclocheilichthys apogon (Valenciennes 1842)    a-, without; pogon, beard, referring to absence of barbels

Cyclocheilichthys armatus (Valenciennes 1842)    armed with a weapon, referring to strongly serrated osseus dorsal fin spine

Cyclocheilichthys heteronema (Bleeker 1854)    heteros, different; nema, thread, referring to how barbels are divided into long fringes, different from all other species of Barbus (original genus) known at the time

Cyclocheilichthys janthochir (Bleeker 1854)    ianthus (with Latin “i” replaced by Roman “j”), violet; cheiros, hand, referring to color of pectoral fin

Cyclocheilichthys lagleri Sontirat 1985    in honor of Karl F. Lagler (1912-1985), for his “intensive” collection of Thai fishes and his work advancing Thai ichthyology

Cyclocheilichthys repasson (Bleeker 1853)    local vernacular in Lampong Province, Sumatra (type locality)

Cyclocheilichthys schoppeae Cervancia & Kottelat 2007    in honor of Sabine Schoppe, for her “lasting help and support to the studies and research of the first author”

Cyclocheilichthys sinensis Bleeker 1879    ensis, suffix denoting place: Sinica (China), where it is endemic

Cyclocheilos Bleeker 1859    cyclo-, round or circular, cheilos, lips, allusion not specified, possibly referring to continuous lips forming round, sucker-like shape when open

Cyclocheilos enoplos (Bleeker 1849)    armed, referring to very strong and serrated osseus dorsal fin spine

Cyclocheilos furcatus Sontirat 1985    forked, referring to bifurcated (and multifurcated) lateral-line tubes in half-grown and adult fish, a character shared with C. enoplos

Cyprinion Heckel 1843    diminutive of Cyprinus, carp, possibly referring to the similarity of C. macrostomum to that of juvenile Cyprinus carpio

Cyprinion acinaces acinaces Banister & Clarke 1977    scimitar, referring to curve of the mouth

Cyprinion acinaces hijazi Krupp 1983    of Hijaz (also spelled Hejaz), western region of Saudi Arabia, where it occurs in Red Sea tributaries

Cyprinion kais Heckel 1843    local name for this fish in Aleppo, Syria (one of the type localities), possibly from the Neo-Syrian kayis, meaning little (reaches 25 cm TL)

Cyprinion macrostomum Heckel 1843    macro-, long; stomus, mouth, referring to large, transverse mouth (“ore transverso, magno”)

Cyprinion mhalense Alkahem & Behnke 1983    ensis, suffix denoting place: Wadi (Valley) Al Mhaleh, southeast Abha City, Saudi Arabia, type locality

Cyprinion microphthalmum (Day 1880)    micro-, small; opthalmus, eye, referring to small eyes, diameter 5½ in length of head, two diameters from end of snout, and 2½ apart

Cyprinion milesi (Day 1880)    in honor of “Col. Miles,” probably Samuel Barrett Miles (1838-1914), British explorer and political agent, who provided type from Afghanistan

Cyprinion semiplotum (McClelland 1839)    semi-, half; plotus, swimmer or floater, allusion not evident, perhaps referring to the observation that larger specimens rise near the surface during the evening and, presumably, are easy to catch with a cast net

Cyprinion tenuiradius Heckel 1847    tenuis, thin; radius, ray, referring to much thinner dorsal spine compared to C. macrostomum

Cyprinion watsoni (Day 1872)    in honor of H. E. Watson, who “largely assisted” Day in collecting natural history specimens on Sind Hills, Pakistan

Cyprinus Linnaeus 1758    kyprinos, Greek for carp, possibly derived from Kypris, also known as Aphrodite, goddess of love, referring to fecundity of C. carpio

Subgenus Cyprinus

Cyprinus acutidorsalis Wang 1979    acutus, sharp; dorsalis, dorsal fin, referring to “triangularly projected” anterior part of dorsal fin [this name may be a junior primary homonym of C. acutidorsalis Chen & Hwang 1977, described from a different part of China and likely representing a different species; more research is needed to confirm the validity of either taxon and the availability of either name]

Cyprinus barbatus Chen & Hwang 1977    bearded or barbeled; described as a subspecies of C. pellegrini, referring to small, minute barbels, compared to absence of barbels on nominate form

Cyprinus carpio Linnaeus 1758    latinization of the Old French carpe

Cyprinus chilia Wu, Yang, Yue & Huang 1963    ia, belonging to: Qilu Lake, Tonghai County, Yunnan Province, China, type locality (“chilu” is the approximate phonetic spelling of “Qilu”)

Cyprinus dai (Nguyen & Doan 1969)    of River Song Da (Da River), between Lai Chau and Hoa Binh, Viêt Nam, type locality

Cyprinus daliensis Chen & Hwang 1977    ensis, suffix denoting place: Dali Erhai Lake, Yunnan Province, China, type locality

Cyprinus exophthalmus Mai 1978    ex, presumably rise up or come out; ophthalmos, eye, referring to large eyes, larger than C. multitaeniata

Cyprinus hyperdorsalis Nguyen 1991    hyper, very; dorsalis, of the back, referring to exceedingly high arched back

Cyprinus ilishaestomus Chen & Huang 1977    Ilisha, a genus of pristigasterid fishes (Clupeiformes); stomus, mouth, referring to Ilisha-like terminal mouth

Cyprinus intha Annandale 1918    Intha, referring to Tibeto-Burman ethnic group living around Inlé Lake (Myanmar), type locality

Cyprinus longipectoralis Chen & Hwang 1977    longus, long; pectoralis, pectoral, referring to long pectoral fins, which reach or extend beyond base of ventral fins

Cyprinus megalophthalmus Wu, Yang, Yue & Huang 1963    mega-, large; ophthalmus, eye, referring to eye diameter larger than mouth diameter

Cyprinus melanes Mai 1978    melanos, black; proposed as a subspecies of Carassioides cantonensis (=acuminatus), referring to its blacker coloration

Cyprinus multitaeniatus Pellegrin & Chevey 1936    multi-, many; taeniata, banded, referring to ~8 dark, parallel, longitudinal lines on body

Cyprinus pellegrini Tchang 1933    in honor of Jacques Pellegrin (1873-1944), “assistant professor,” Muséum national d’histoire naturelle (Paris) [and author of several Chinese cyprinids]

Cyprinus qionghaiensis Liu 1981    ensis, suffix denoting place: Qionghai Lake, Xichang Xian, Sichuan Province, China, type locality

Cyprinus quidatensis Nguyen, Le, Le & Nguyen 1999    ensis, suffix denoting place: Qui Dat, Minh Hoa, Quang Binh, Viêt Nam, type locality

Cyprinus rubrofuscus Lacepède 1803    rubra-, red; fuscus, dusky or dark, referring to golden-brown coloration

Cyprinus yunnanensis Tchang 1933    ensis, suffix denoting place: Yunnan Province, China, where it is endemic

Subgenus Mesocyprinus Fang 1936    meso-, middle, allusion unclear, perhaps referring to Fang’s belief that pharyngeal bone and teeth structure of C. micristius indicated a “more advanced stage of evolution,” thereby representing an intermediate or middle form between Cyprinus and other cyprinids

Cyprinus fuxianensis Yang et al. 1977    ensis, suffix denoting place: Fuxian Lake, Yunnan Province, China, type locality [note: original description gives authorship as “Yang et al.” with no explanation of who the other authors might be]

Cyprinus longzhouensis Yang & Hwang 1977    ensis, suffix denoting place: Lungzhou, Guangxi, China, type locality

Cyprinus micristius Regan 1906    micro-, small; istios, sail referring to smaller number of dorsal-fin rays compared to C. carpio

Cyprinus yilongensis Yang et al. 1977    ensis, suffix denoting place: Yi-lung (Yulong) Lake, Shiping, Yunnan Province, China, where species (now extinct) was endemic [note: original description gives authorship as “Yang et al.” with no explanation of who the other authors might be]

Dawkinsia Pethiyagoda, Meegaskumbura & Maduwage 2012    ia, belonging to: ethologist and evolutionary biologist Richard Dawkins, for his contribution to the public understanding of science

Dawkinsia arulius (Jerdon 1849)    latinization of aruli, Kannada vernacular for this species

Dawkinsia assimilis (Jerdon 1849)    similar, i.e., “very closely allied” to D. filamentosa

Dawkinsia exclamatio (Pethiyagoda & Kottelat 2005)    referring to color pattern, which, if viewed “snout-down,” resembles an exclamation mark (!)

Dawkinsia filamentosa (Valenciennes 1844)    filamentous, referring to branched dorsal-fin rays prolonged into filament-like extensions (in adult males only)

Dawkinsia rohani (Rema Devi, Indra & Knight 2010)    in honor of Rohan Pethiyagoda, for his extensive work on the freshwater fishes of India and Sri Lanka

Dawkinsia rubrotinctus (Jerdon 1849)    rubro-, red; tinctus, colored, referring to series of bright red spots on sides

Dawkinsia singhala (Duncker 1912)    ancient name of Sri Lanka, where it is endemic

Dawkinsia srilankensis (Senanayake 1985)    ensis, suffix denoting place: Sri Lanka, where it is endemic

Dawkinsia tambraparniei (Silas 1954)    of the Tambraparni watershed, Tinnevelly District, Madras State, India, only known area of occurrence

Desmopuntius Kottelat 2013    desmotes, prisoner, referring to color pattern made of 4-6 bars (at least in juveniles), which distinguishes it from other genera formerly placed in Puntius in Southeast Asia

Desmopuntius endecanalis (Roberts 1989)    en-, within; decem, ten; analis, anal, referring to 8 (rarely 7) branched anal-fin rays, whereas most Puntius (former genus) have 5 or 6

Desmopuntius foerschi (Kottelat 1982)    in honor of physician and aquarist Walter Foersch, who helped collect type

Desmopuntius gemellus (Kottelat 1996)    twin, referring to similarity to D. johorensis

Desmopuntius hexazona (Weber & de Beaufort 1912)    hexa-, six; zonatus, banded, referring to six black, transverse bands on body

Desmopuntius johorensis (Duncker 1904)    –ensis, suffix denoting place: Johore, Malaysia, type locality

Desmopuntius pentazona (Boulenger 1894)    penta-, five; zonus, belt or girdle, referring to five blackish bands that encircle body

Desmopuntius rhomboocellatus (Koumans 1940)    rhombus, diamond-shaped; ocellatus, having little eyes, referring to how predorsal, dorsal and anal bars expand to form “ocellate rhombi”

Desmopuntius trifasciatus (Kottelat 1996)    tri-, three; fasciatus, banded, referring to 3-4 bands on sides of juveniles, becoming 3-4 stripes on sides of adults

Diplocheilichthys Bleeker 1859    ichthys, fish, with diplo– (twofold) and cheilus (lip), presumably referring to inner transverse fold on upper lip of D. pleurotaenia, which effectively divides lip in half

Diplocheilichthys jentinkii (Popta 1904)    in honor of Fredericus Anna Jentink (1844-1913), curator at Rijksmuseum van Natuurlijke Historie (now Naturalis) in Leiden, Netherlands, and editor of its journal (in which description appeared), for making the museum’s specimens available to Popta

Diplocheilichthys pleurotaenia (Bleeker 1855)    pleuro-, side; taenia, ribbon, referring to lateral stripe from gill opening to end of caudal peduncle (usually more distinct in younger specimens, sometimes absent in adults)

Diptychus Steindachner 1866    di-, two; ptychos, fold, referring to lower jaw with a cutting anterior edge covered with a horny sheath, behind which is a distinct, uninterrupted labial fold, continuous with upper lip

Diptychus maculatus Steindachner 1866    spotted, referring to two rows of dark speckles on abdomen and lower portion of head

Diptychus sewerzowi Kessler 1872    in honor of zoologist Nikolai A. Severtzov (also spelled Severzov, Sewerzoff, 1827-1885), who collected type

Discherodontus Rainboth 1989    di-, two; scheros, in a line; dontus, tooth, referring to two rows of pharyngeal teeth, a character unique among barbins

Discherodontus ashmeadi (Fowler 1937)    in honor of Charles C. Ashmead, an early local contributor to the fish collection at the Academy of Natural Sciences of Philadelphia

Discherodontus colemani (Fowler 1937)    in honor of Waldburg Coleman, early contributor to the fish collection at the Academy of Natural Sciences of Philadelphia

Discherodontus halei (Duncker 1904)    in honor of Abraham Hale, Museum Committee chair, Selangor State Museum (Malaysia), for “sympathetic kindness” when Duncker took over the museum’s “small, ill-kept collection of stuffed fishes” (translations)

Discherodontus parvus (Wu & Lin 1977)    small, presumably referring to small size, 50-52 mm SL

Discherodontus schroederi (Smith 1945)    in honor of William C. Schroeder (1895-1977), associate curator of fishes, Museum of Comparative Zoology, Harvard University, for making available a collection of Thai fishes from 1937

Discherodontus somphongsi (Benl & Klausewitz 1962)    in honor of Mr. Somphongs, Bangkok, presumably of Somphongs Aquarium Company, who collected the first specimens and provided the authors with living and preserved material

Discocheilus Zhang 1997    discus, disk; cheilus, lip, referring to lower lip modified into an adhesive or suctorial disk (replacement name for Discolabeo Chen & Lan 1992, preoccupied by Discolabeo Fowler 1937)

Discocheilus multilepis (Wang & Li 1994)    multi-, many; lepis, scale, having more scales than D. wui

Discocheilus wui (Chen & Lan 1992)    in honor of the late Xian-Wen Wu, who dedicated his research to this fish and for his many publications

Discogobio Lin 1931    discus, disk, referring to lower lip modified into an adhesive or suctorial disk; gobio, reflecting original placement in subfamily Gobioninae

Discogobio antethoracalis Zheng & Zhou 2008    ante, anterior; thoracalis, thoracic, referring to anterior position of pectoral fins

Discogobio bismargaritus Chu, Cui & Zhou 1993    bis, twice or double; margaritus, pearl, referring to pair of pearl organs on snout

Discogobio brachyphysallidos Huang 1989    brachys, short; physallidos, bladder, referring to shorter air bladder (shorter than length of head) compared to most other Chinese congeners

Discogobio caobangi Nguyen 2001    of Cao Bàng Province, Viêt Nam, where it is endemic

Discogobio dienbieni Nguyen 2001    of Diên Biên, Lai Chau Province, Viêt Nam, where it is endemic

Discogobio elongatus Huang 1989     elongated, referring to elongate body, with a length-to-height ratio of 6.0-7.5

Discogobio laticeps Chu, Cui & Zhou 1993    latus, wide; ceps, head, referring to head that is wider than deep

Discogobio longibarbatus Wu 1977    longus, long; barbatus, barbled, referring to long barbels, reaching below or beyond middle of eyes

Discogobio macrophysallidos Huang 1989    macro-, long; physallidos, bladder, referring to longer air bladder (equal to or longer than length of head) compared to most other Chinese congeners

Discogobio microstoma (Mai 1978)    micro-, small; stoma, mouth, referring to small “sucker mouth” (translation) or adhesive disc formed by modified upper lip

Discogobio multilineatus Cui, Zhou & Lan 1993    multi-, many; lineatus, lined, referring to 5-6 black slender longitudinal lines on body from posterior of head to caudal-fin base

Discogobio pacboensis Nguyen 2001    ensis, suffix denoting place: Pac Bo, Cao Bang Province, Viêt Nam, where it is endemic

Discogobio poneventralis Zheng & Zhou 2008    pone, posterior; ventralis, ventral, referring to posterior position of pelvic fins

Discogobio propeanalis Zheng & Zhou 2008    prope, close; analis, anal, referring to position of pelvic fins, which are closer to anal-fin origin

Discogobio tetrabarbatus Lin 1931    tetra-, four; barbatus, bearded, referring to two pairs of small barbels on lower jaw

Discogobio yunnanensis (Regan 1907)    ensis, suffix denoting place: Yunnan Province, China, type locality

Discolabeo Fowler 1937    discus, disk, like a Labeo with a disk-like mandible

Discolabeo wuluoheensis Li, Lu & Mao 1996    ensis, suffix denoting place: Wuluo He (=River), Yunnan Province, China, type locality

Eechathalakenda Menon 1999    etymology not explained, presumably a variant of eetilakanda, Malayam name for this fish

Eechathalakenda ophicephala (Raj 1941)    ophis, snake; cephalus, head, referring to elongate shape, similar in appearance to snakeheads (Perciformes: Channidae)

Eirmotus Schultz 1959    otus, adjectival suffix: eirmos, series, referring to “series of sensory pores on the head”

Eirmotus furvus Tan & Kottelat 2008    dark black or dusky, referring to overall dusky appearance

Eirmotus insignis Tan & Kottelat 2008    worth distinction, with distinctive signs, or brilliant, referring to unique (in genus) black marks on dorsal fin

Eirmotus isthmus Tan & Kottelat 2008    narrow passage or neck of land, referring to narrow black bars on body

Eirmotus octozona Schultz 1959    octo-, eight; zona, band, referring to eight blackish bars on head and body

Enteromius Cope 1867    intestinal, referring to short alimentary canal of E. potamogalis

Enteromius ablabes (Bleeker 1863)    harmless, referring to unserrated dorsal fin spine

Enteromius aboinensis (Boulenger 1911)    ensis, suffix denoting place: headwaters of Aboina River, southern Nigeria, type locality

Enteromius afrohamiltoni (Crass 1960)    afro-, African, in honor of Lt.-Col. J. Stevenson Hamilton, warden, Kruger National Park, who collected type [replacement name for Barbus hamiltoni Gilchrist & Thompson 1913, preoccupied by B. hamiltonii (Jerdon 1849) (=Bangana ariza) of India, i.e., “the African hamiltoni”]

Enteromius afrovernayi (Nichols & Boulton 1927)    afro-, African; replacement name for Barbus vernayi Nichols & Boulton 1927, preoccupied by B. (=Hypsibarbus) vernayi Norman 1925 of China, i.e., “the African vernayi,” in honor of Arthur S. Vernay (1877-1960), art and antiques dealer, big game hunter and naturalist-explorer, who funded and led an American Museum of Natural History expedition to Angola

Enteromius aliciae (Bigorne & Lévêque 1993)    in honor of the second author’s daughter (presumably Alice)

Enteromius aloyi (Roman 1971)    in honor of Isidro Aloy (relationship to author not specified), for “his assistance at all times” (translation)

Enteromius altidorsalis (Boulenger 1908)    altus, high; dorsalis, dorsal, referring to high anterior rays of dorsal fin

Enteromius amanpoae (Lambert 1961)    of Amanpoa River, Ponthierville Territory, Congo, type locality

Enteromius amatolicus (Skelton 1990)    icus, belonging to: Amatola Mountains (Eastern Cape Province, South Africa), which give rise to Isidenge River, type locality

Enteromius anema (Boulenger 1903)    a-, without; nema, thread, presumably referring to lack of barbels

Enteromius annectens (Gilchrist & Thompson 1917)    linking or joining, perhaps referring to how it “closely resembles” E. lineomaculatus and Barbus werneri (=E. stigmatopygus), thus serving as a “link” between the two

Enteromius anniae (Lévêque 1983)    in honor of Lévêque’s wife Annie (Christian Lévêque, pers. comm., matronym not identified in description)

Enteromius anoplus (Weber 1897)    unarmed, allusion not explained, perhaps referring to unserrated last dorsal-fin ray

Enteromius ansorgii (Boulenger 1904)    in honor of explorer William John Ansorge (1850-1913), who collected type

Enteromius apleurogramma (Boulenger 1911)    a-, without; pleuro-, side; gramme, line, referring to “no lateral line” (actually an incomplete or reduced lateral line, with 0-5 pore-bearing scales)

Enteromius arambourgi (Pellegrin 1935)    probably in honor of vertebrate paleontologist Camille Arambourg (1885-1970), who conducted extensive field work in North Africa (including Ethiopia, where this barb is endemic)

Enteromius arcislongae (Keilhack 1908)     etymology not explained but here is a guess: arcis, arc or bow, referring to downward arc of lateral line seen in many small Enteromius; long, referring to greater number of lateral line scales compared to E. trispilopleura, which would make downward arc of lateral line longer

Enteromius argenteus (Günther 1868)    silvery, referring to “uniform silvery” coloration

Enteromius aspilus (Boulenger 1907)    a-, without; spilos, spot, referring to lack of markings

Enteromius atakorensis (Daget 1957)    ensis, suffix denoting place: a creek in Atakora Mountains, Republic of Benin, type locality

Enteromius atkinsoni (Bailey 1969)    in honor of the late Maurice Atkinson, Lake Victoria and Tanzanian Fisheries Service, “who had a wide interest in the biology and correct identification of East African fishes, and whose contributions in the realm of fisheries development and training, will long be valued by his colleagues and students alike”

Enteromius atromaculatus (Nichols & Griscom 1917)    atro-, black; maculatus, spotted, referring to 3-7 black spots on each side, connected by a dark lateral streak

Enteromius bagbwensis (Norman 1932)    ensis, suffix denoting place: Bagbwe River, Sierra Leone, type locality

Enteromius barnardi (Jubb 1965)    in honor of Keppel Harcourt Barnard (1887-1964), South African Museum, for his “remarkable contributions” to the taxonomy of South African fishes

Enteromius barotseensis (Pellegrin 1920)    ensis, suffix denoting place: Barotsés, northwestern Rhodesia, type locality

Enteromius baudoni (Boulenger 1918)    in honor of in honor of French colonial administrator Alfred Baudon (1875-1932), who sent to the British Museum (Natural History) a collection of fishes from the Shari River, including type of this one

Enteromius bawkuensis (Hopson 1965)    ensis, suffix denoting place: White Volta and its tributaries near Bawku, northern Ghana, only known distribution

Enteromius bifrenatus (Fowler 1935)    bi-, two; frenatus, brindled, referring to two distinct black lateral stripes beginning on snout and running through eye and gill cover and down body

Enteromius bigornei (Lévêque, Teugels & Thys van den Audenaerde 1988)    in honor of friend and colleague Rémy Bigorne, ichthyologist, ORSTOM (Office de la Recherche Scientifique et Technique d’Outre-Mer)

Enteromius boboi (Schultz 1942)    in honor of one of the local men who helped William M. Munn, director of the National Zoological Park (Washington, D.C.), collect fishes in Liberia

Enteromius bourdariei (Pellegrin 1928)    in honor of Paul Bourdarie (1864-1950), co-founder and permanent secretary of l’Académie des Sciences Coloniales (now l’Académie des sciences d’outre-mer), a learned society dedicated to the geography and history of Africa, Latin America, Asia and Oceania

Enteromius brachygramma (Boulenger 1915)    brachys, short; gramme, line, presumably referring to incomplete lateral line and/or “poorly defined” (translation) blackish lateral band

Enteromius brazzai (Pellegrin 1901)    in honor of Franco-Italian explorer Pierre Savorgnan de Brazza (1852-1905), who collected type

Enteromius breviceps (Trewavas 1936)    brevis, short; ceps, head, referring to smaller head compared to E. unitaeniatus

Enteromius brevidorsalis (Boulenger 1915)    brevis, short; dorsalis, dorsal, referring to shorter second dorsal ray compared to other barbs with a single pair of barbels described in same paper

Enteromius brevilateralis (Poll 1967)    brevis, short; lateralis, of the side, referring to shorter lateral line compared to E. kessleri

Enteromius brevipinnis (Jubb 1966)    brevis, short; pinna, fin, referring to very short fins on type specimens from Merite River (probably caused by environmental factors and not typical of most populations)

Enteromius brichardi (Poll & Lambert 1959)    in honor of aquarium-fish exporter Pierre Brichard (1921-1990), who collected type

Enteromius cadenati (Daget 1962)    in honor of ichthyologist Jean Cadenat (1908-1992), Director, Marine Biological Section of the Institute Français d’Afrique Noire, which published Daget’s monograph, and who collected one specimen in 1947

Enteromius callipterus (Boulenger 1907)    calli-, beautiful; pterus, fin, referring to orange (basal) and white (distal) dorsal fin, with a large deep spot between last simple ray and branched third ray

Enteromius camptacanthus (Bleeker 1863)    camptos, bent or curved; acanthus, thorn, referring to more flexible dorsal fin spine compared to other Puntius (genus in which it was described)

Enteromius candens (Nichols & Griscom 1917)    shining or gleaming, presumably referring to its being a “very distinct and beautiful little species”

Enteromius carcharhinoides (Stiassny 1991)    oides, having the form of: carcharias, man-eating shark (from karcharos, jagged, referring to a shark’s rasp-like skin), referring to barb’s large and strongly falcate dorsal fin, which gives it a shark-like appearance

Enteromius carens (Boulenger 1912)    from carere, to be without or devoid of, referring to absence of barbels and any indication of a lateral line

Enteromius castrasibutum (Fowler 1936)    castrum, fort, referring to Fort Sibut, Ubangi-Shari (now Central African Republic), type locality

Enteromius catenarius (Poll & Lambert 1959)    chained, allusion not evident, perhaps referring to interrupted black longitudinal band composed of 3-5 elongated spots, and/or scales on upper part of body largely (or completely) bordered in brown, either attribute forming a chain-like effect

Enteromius caudosignatus (Poll 1967)    caudo-, tail; signatus, marked, referring to large oval black spot at base of tail

Enteromius cercops (Whitehead 1960)    cerco-, tail; ops, face, after the Cercopes, Heracles’ mischievous tormentors, referring to long barbels

Enteromius chicapaensis (Poll 1967)    ensis, suffix denoting place: Alto Chicapa, marsh along River Coca, Angola, type locality

Enteromius chiumbeensis (Pellegrin 1936)    ensis, suffix denoting place: Kasai River at Chiumbé, Angola, type locality

Enteromius chlorotaenia (Boulenger 1911)    chloros, green; taenia, band, referring to dark green (black in spirits) lateral stripe

Enteromius choloensis (Norman 1925)    ensis, suffix denoting place: Cholo, Nyasaland (Malawi), type locality

Enteromius citrinus (Boulenger 1920)    like a citron (a lemon-like fruit), referring to lemon-yellow coloration of body and fins

Enteromius clauseni (Thys van den Audenaerde 1976)    in honor of Danish ichthyologist H. Stenholt Clausen, who collected type

Enteromius collarti (Poll 1945)     in honor of entomologist Albert Collart (1899-1993), who collected type

Enteromius condei (Mahnert & Géry 1982)    in honor of zoologist Bruno Condé (1920-2004), director of l’Aquarium de Nancy, who helped collect type

Enteromius deguidei (Matthes 1964)    in honor of R. Deguide, Musée Royal de l’Afrique Centrale, for his invaluable assistance during Matthes’ research in the Ikela region of the Congo

Enteromius deserti (Pellegrin 1909)    presumably “of the desert,” referring to occurrence in Algerian Sahara, which Pellegrin found to be quite remarkable

Enteromius devosi (Banyankimbona, Vreven & Snoeks 2012)    in honor of the late Luc De Vos (1957-2003), “outstanding Belgian ichthyologist and dear colleague and friend,” whose “remarkable collecting efforts and scientific determination” contributed to the knowledge of Malagarazi fishes, and who was the first to recognize this species as new to science

Enteromius dialonensis (Daget 1962)    ensis, suffix denoting place: northeastern Fouta Dialon (also spelled Djallon), highland region in the center of Guinea, where it occurs

Enteromius diamouanganai (Teugels & Mamonekene 1992)    in honor of Jean Diamouangana, UNESCO National Project Director in Mayombe, Congo, which supported the authors’ work

Enteromius ditinensis (Daget 1962)    ensis, suffix denoting place: Ditinn, Lower Guinea, type locality

Enteromius dorsolineatus (Trewavas 1936)    dorso-, dorsal or back; lineatus, lined, referring to dark stripe before dorsal fin

Enteromius eburneensis (Poll 1941)    ensis, suffix denoting place: eburn, ivory, referring to type locality in the Ivory Coast

Enteromius erythrozonus (Poll & Lambert 1959)    erythros, reddish; zonus, band, referring to broad gold-red longitudinal band

Enteromius eutaenia (Boulenger 1904)    eu-, very; taenia, band, referring to “sharply defined” black lateral band passing through eyes and extending to caudal fin

Enteromius evansi (Fowler 1930)    in honor of J. R. Evans, who accompanied Fowler on the Gray African Expedition in Angola (during which type was collected) and “whose knowledge of the country and its languages was invaluable to the success of the expedition”

Enteromius fasciolatus (Günther 1868)    banded, referring to ~12 narrow blackish-blue vertical streaks on sides

Enteromius foutensis (Lévêque, Teugels & Thys van den Audenaerde 1988)    ensis, suffix denoting place: Fouta Djallon, highland region in the middle of Guinea, West Africa, where it appears to be endemic

Enteromius greenwoodi (Poll 1967)    in honor of Peter Humphry Greenwood (1927-1995), author of many papers on African fishes in general and Barbus in particular

Enteromius guildi (Loiselle 1973)    in honor of Paul D. Guild, friend and Peace Corps colleague for three years in the Republic of Togo

Enteromius guineensis (Pellegrin 1913)    ensis, suffix denoting place: Guinea, where it is endemic

Enteromius guirali (Thominot 1886)    in honor of explorer and naturalist Léon Guiral (1858-1885), who collected type in Zaire (now Democratic Republic of the Congo) in 1885 (and who died, presumably of yellow fever, shortly thereafter)

Enteromius gurneyi (Günther 1868)    in honor of John Henry Gurney (1819-1890), financier and ornithologist, through whose “mediation” Günther received numerous specimens from Port Natal (now Durban, South Africa)

Enteromius haasianus (David 1936)    anus, belonging to: malacologist Fritz Haas (1886-1969), who collected type

Enteromius holotaenia (Boulenger 1904)    holo-, entire; taenia, band, referring to black band running entire length of body, from snout to caudal fin base

Enteromius huguenyi (Bigorne & Lévêque 1993)    in honor of friend and colleague Bernard Hugueny, Institut de Recherche pour le Développement (IRD)

Enteromius hulstaerti (Poll 1945)    in honor of entomologist R. P. Hulstaert, who collected type

Enteromius humeralis (Boulenger 1902)    of the shoulder, presumably referring to black vertical bar behind gill opening

Enteromius humilis (Boulenger 1902)    low or humble, allusion not evident, perhaps not as colorful or attractive as some or most of the other Abyssianian Barbus Boulenger described in the same paper

Enteromius inaequalis (Lévêque, Teugels & Thys van den Audenaerde 1988)    “inégal, difficile à reconnaître” (unequal [i.e., no others are quite like it], difficult to recognize), meaning unclear, possibly referring to previous misidentification as E. boboi

Enteromius innocens (Pfeffer 1896)    harmless, presumably referring to “weak and pliable” (translation) dorsal-fin ray

Enteromius jacksoni (Günther 1889)    in honor of Frederick John Jackson (1859-1929), English administrator, explorer and ornithologist, who “obtained” type

Enteromius jae (Boulenger 1903)    of the Ja River, southern Cameroon, type locality

Enteromius janssensi (Poll 1976)   dedicated to the memory of entomologist André Janssens (1906-1954), who participated in a large-scale faunal survey (1946-1949) of Upemba National Park (Democratic Republic of the Congo), type locality

Enteromius kamolondoensis (Poll 1938)    ensis, suffix denoting place: Kamolondo, a plain that crosses the Lualaba River downstream of Bukama, Democratic Republic of the Congo, type locality

Enteromius kerstenii (Peters 1868)    in honor of Otto Kersten (1839-1900), early explorer of Mt. Kilimanjaro, who sent a small collection of fishes to Peters, including type of this one

Enteromius kessleri (Steindachner 1866)    patronym not identified but probably in honor of German-Russian zoologist Karl Federovich Kessler (1815-1881), who described at least two cyprinids by 1866 (and many more thereafter)

Enteromius kissiensis (Daget 1954)    ensis, suffix denoting place: Kissidougon, Guinea, type locality, where Kissi is the largest ethnic group

Enteromius kuiluensis (Pellegrin 1930)    ensis, suffix denoting place: Kouilou River, Gabon, type locality

Enteromius lamani (Lönnberg & Rendahl 1920)    in honor of Swedish missionary and ethnographer Karl Edvard Laman (1867-1944), who sent a small collection of fishes from the lower Congo to the Riksmuseet Natural History Museum in Stockholm, including type of this one

Enteromius laticeps (Pfeffer 1889)    latus, wide; ceps, head, referring to very wide, dorsally flattened head

Enteromius lauzannei (Lévêque & Paugy 1982)    in honor of friend and colleague Laurent Lauzanne

Enteromius leonensis (Boulenger 1915)    ensis, suffix denoting place: Maka, Sierra Leone, type locality

Enteromius liberiensis (Steindachner 1894)    ensis, suffix denoting place: Grand Cape Mount and Robertsport, Liberia, type locality

Enteromius lineomaculatus (Boulenger 1903)    lineo-, line; maculatus, spotted, referring to series of irregular black spots on sides, more or less connected by a dark lateral streak

Enteromius litamba (Keilhack 1908)    local name for this barb at Lake Malawi

Enteromius lornae (Ricardo-Bertram 1943)    in honor of Lorna Brown, wife of English aristocrat Sir Stewart Gore-Brown, residents of Shiwa Ngandu, an English-style country house and estate in Zambia, who helped and advised the author during her expedition

Enteromius loveridgii (Boulenger 1916)    in honor of herpetologist Arthur Loveridge (1891-1980), Curator of the Nairobi Museum (barb is endemic to Kenya)

Enteromius lufukiensis (Boulenger 1917)    ensis, suffix denoting place: Lufuko River at Pala, Lake Tanganyika, type locality

Enteromius luikae (Ricardo 1939)    of Luika Pool, below falls of Luika River (Tanzania), type locality

Enteromius lujae (Boulenger 1913)    in honor of botanist and entomologist Edouard Luja (1875-1953), resident of Kasai, Zaire (type locality), who collected type

Enteromius lukindae (Boulenger 1915)    of the Lukinda River drainage, Democratic Republic of the Congo, type locality

Enteromius lukusiensis (David & Poll 1937)    ensis, suffix denoting place: Lukusi River, Democratic Republic of the Congo, type locality

Enteromius luluae (Fowler 1930)    of the Lulua River, Democratic Republic of the Congo, type locality

Enteromius machadoi (Poll 1967)    in honor of António de Barros Machado (1912-2002), “distinguished” (translation) zoologist of the Musée de Dundo (type locality in Dundo, Angola)

Enteromius macinensis (Daget 1954)    ensis, suffix denoting place: Macina (Inner Niger Delta), where it is endemic

Enteromius macrops (Boulenger 1911)    macro-, long; ops, eye, referring to much larger eye compared to E. ablabes

Enteromius macrotaenia (Worthington 1933)    macro-, long; taenia, band, referring to broad, black stripe from tip of snout to base of caudal fin

Enteromius magdalenae (Boulenger 1906)    in honor of Magdalene (or Magdalena) Milchin, wife of zoologist of E. A. Milchin, whom Boulenger honored in the same paper for his service to African ichthyology

Enteromius manicensis (Pellegrin 1919)    ensis, suffix denoting place: Manica, Mozambique, type locality

Enteromius marmoratus (David & Poll 1937)    marbled or mottled, probably referring to scales edged with black and/or black stripe on the sides divided into smaller vertical bands

Enteromius martorelli (Roman 1971)    in honor of Fernando Martorell (relationship to author not specified), as a “token of gratitude” (translation)

Enteromius mattozi (Guimarães 1884)    in honor of Ferdinand dos Santos Mattozo, zoology professor, l’École Polytechnique de Lisbonne

Enteromius mediosquamatus (Poll 1967)    medio-, middle; squamatus, scaled, probably referring to taller size of lateral line scales compared to adjacent scales

Enteromius melanotaenia (Stiassny 1991)   melanos, black; taeniata, banded, referring to broad, ribbonlike black lateral stripe

Enteromius mimus (Boulenger 1912)    imitator, referring to “striking resemblance” to E. neglectus

Enteromius miolepis (Boulenger 1902)    mio-, less or small; lepis, scale, presumably referring to fewer scale rows below lateral line compared to E. kessleri

Enteromius mocoensis (Trewavas 1936)    ensis, suffix denoting place: a brook at Mt. Moco, Cuvo River system, Angola, type locality

Enteromius mohasicus (Pappenheim 1914)    icus, belonging to: Lake Mohasi, Rwanda, type locality

Enteromius motebensis (Steindachner 1894)    ensis, suffix denoting place: Motebe River, Marico District, South Africa, type locality

Enteromius multilineatus (Worthington 1933)    multi-, many; lineatus, lined, referring to series of three broken bands above lateral stripe and two below

Enteromius musumbi (Boulenger 1910)    native name for this barb in Angola

Enteromius neefi (Greenwood 1962)    in honor of neef, Afrikaans word for nephew, a humorous acknowledgment to Graham Bell-Cross, who collected type and often called Greenwood oom, Afrikaans word or uncle [Paul Skelton, pers. comm.]

Enteromius neglectus (Boulenger 1903)    overlooked; Boulenger had initially regarded the species as the young of E. perince

Enteromius neumayeri (Fischer 1884)    in honor of George von Neumayer (1826-1909), director of German Marine Observatory and vice-chairman of the Geographical Society in Hamburg

Enteromius nigeriensis (Boulenger 1903)    ensis, suffix denoting place: Niger River Delta, type locality

Enteromius nigrifilis (Nichols 1928)    niger, black; filum, thread, referring to black “thread-like” stripe on side from shoulder to base of caudal fin

Enteromius nigroluteus (Pellegrin 1930)    niger, black; luteus, yellow, referring to yellowish back, sulfur-yellow snout, and broad black band along the sides

Enteromius niokoloensis (Daget 1959)    ensis, suffix denoting place: Niokolo-Koba National Park, Senegal, type locality

Enteromius nounensis (Van den Bergh & Teugels 1998)    ensis, suffix denoting place: Noun River, Cameroon, type locality

Enteromius nyanzae (Whitehead 1960)    of Nyanza Province, Kenya, type locality

Enteromius okae (Fowler 1949)    of Oka, Democratic Republic of the Congo, type locality

Enteromius oligogrammus (David 1937)    oligo-, few; gramme, line, referring to incomplete lateral line, just 4-13 scales

Enteromius olivaceus (Seegers 1996)    referring to olive-green coloration

Enteromius owenae (Ricardo-Bertram 1943)    in honor of Miss R. J. Owen, who studied fishes with the author in Lake Rukwa (Tanzania) and the Bangweulu Region of Zambia

Enteromius pallidus (Smith 1841)    pale, presumably referring to silvery sides and ventral surface in females and non-breeding males

Enteromius paludinosus (Peters 1852)    swampy or marshy, presumably referring to barb’s preference for well-vegetated swamp- and marsh-like waters

Enteromius papilio (Banister & Bailey 1979)    butterfly, referring to “striking” color pattern (deep brown lateral band on a “pale sandy khaki” body), especially the dark fins

Enteromius parablabes (Daget 1957)    para-, near, referring to similarity to E. ablabes

Enteromius parajae (Van den Bergh & Teugels 1998)    para-, near, referring to close morphological relationship to E. jae

Enteromius pellegrini (Poll 1939)    in honor of ichthyologist Jacques Pellegrin (1873-1944), who described this barb in 1935 but used a preoccupied name (Barbus serrifer trimaculata)

Enteromius perince (Rüppell 1835)    vernacular for this barb in the markets of Cairo, Egypt

Enteromius petchkovskyi (Poll 1967)    in honor of “Monsieur” de Petchkovsky (no forename given), for his help in collecting fishes, presumably in Angola

Enteromius pleurogramma (Boulenger 1902)    pleuro-, side; gramme, line, referring to black line on each side of body

Enteromius pobeguini (Pellegrin 1911)    in honor of Charles Henri Oliver Pobeguin (1856-1951), botanist and colonial administrator in French Africa, who sent type specimen to Muséum d’histoire naturelle de Paris in 1904

Enteromius poechii (Steindachner 1911)    in honor of Austrian anthropologist and ethnologist Rudolf Pöch (1870-1921), who collected type

Enteromius potamogalis Cope 1867    potamos, river; galeus, weasel, presumably alluding to piscivorous Otter Shrew, Potamogale velox, described a year later by Paul Belloni Du Chaillu, who collected type of this barb and included an illustration showing an Otter Shrew on the bank of a river with a large barb in its forepaws in his 1867 book on his African travels

Enteromius prionacanthus (Mahnert & Géry 1982)    prion, saw; acanthus, thorn, referring to strongly ossified last simple ray of dorsal fin, serrated on posterior side

Enteromius profundus (Greenwood 1970)    deep, endemic to the deep waters of Lake Victoria

Enteromius pseudotoppini (Seegers 1996)    pseudo-, false, although this species resembles (and was previously misidentified as) E. toppini, such an appearance is false

Enteromius pumilus (Boulenger 1901)    dwarfish, diminutive or little, referring to small size (type specimens up to 26 mm TL)

Enteromius punctitaeniatus (Daget 1954)    punctus, spot; taeniatus, banded, referring to lateral band formed by a discontinuous series of black spots

Enteromius pygmaeus (Poll & Gosse 1963)    dwarf, referring to small size, up to 25.8 mm TL

Enteromius quadrilineatus (David 1937)    quadri-, four; lineatus, lined, referring to 3-4 black streaks above and below lateral line

Enteromius quadripunctatus (Pfeffer 1896)    quadri-, four; punctatus, spotted, referring to 3-4 small round blackish spots on each side, the last at base of caudal

Enteromius radiatus radiatus (Peters 1853)    rayed, probably referring to radial striations of scales (Peters did not mention this character in his brief original description, but mentioned and illustrated it in an expanded description published in 1868)

Enteromius radiatus aurantiacus (Boulenger 1910)    orange-colored, referring to yellowish lower section tinged with orange, and bright orange basal half or two-thirds of fins

Enteromius raimbaulti (Daget 1962)    in honor of M. (Monsieur) Raimbault (forename not given), inspector, Eaux et Forêts (Waters and Forests), for his role in facilitating Daget’s missions in Guinea and/or help collecting fishes

Enteromius rohani (Pellegrin 1921)    in honor of explorer Jacques de Rohan-Chabot (1889-1958), who collected type

Enteromius roussellei (Ladiges & Voelker 1961)    in honor of Ardo Rousselle, on whose plantation in Angola the type was collected

Enteromius rouxi (Daget 1961)    in honor of Charles Roux, Muséum national d’Histoire naturelle (Paris), who collected type

Enteromius rubrostigma (Poll & Lambert 1964)    rubro-, red; stigma, mark or spot, referring to red spot on opercle

Enteromius salessei (Pellegrin 1908)    in honor of M. (=Monsieur) Salesse, who “greatly facilitated” (translation) the ichthyological research of Dr. Wurtz (see Labeobarbus wurtzi) during the latter’s travels through Senegal and French Guinea

Enteromius sensitivus (Roberts 2010)    sensitive, referring to numerous pitline sensory organelles characteristic of this species

Enteromius serengetiensis (Farm 2000)    ensis, suffix denoting place: Serengeti National Park, Tanzania, type locality

Enteromius sexradiatus (Boulenger 1911)    sex, six; radiatus, rayed, presumably referring to six soft dorsal fin rays

Enteromius seymouri (Tweddle & Skelton 2008)    in honor of the late Tony Seymour, Malawi Government Fisheries Department, close friend and colleague of the first author, for his many years of environmental management and conservation service, and for his long-term commitment to supporting Lake Malawi’s fishermen

Enteromius stanleyi (Poll & Gosse 1974)    in memory of Henry Morton Stanley (1841-1904), “du grand explorateur” of the Congo Basin

Enteromius stauchi (Daget 1967)    in honor of M. A. Stauch, Agent Technique de l’O.R.S.T.O.M. (Office de la Recherche Scientifique et Technique d’Outre-Mer), who collected type

Enteromius stigmasemion (Fowler 1936)    stigma, spot; semeion, banner, referring to black blotch on dorsal fin

Enteromius stigmatopygus (Boulenger 1903)    stigmatus, marked; pygus, rump or buttock, presumably referring to small round black spot at base of anal fin

Enteromius subinensis (Hopson 1965)    ensis, suffix denoting place: Subin River, Prah basin, Juaso, Ghana, type locality

Enteromius sublineatus (Daget 1954)    sub, under; lineatus, lined, referring to how lateral line dips one scale row under dorsal fin

Enteromius sylvaticus (Loiselle & Welcomme 1971)    aticus, belonging to: sylva, forest, referring to forest-stream habitat

Enteromius syntrechalepis (Fowler 1949)    syn-, together; trecha, move; lepis, scale, referring to body scales “converging smaller on tail posteriorly and on caudal peduncle”

Enteromius taeniopleura (Boulenger 1917)    taenio-, band; pleuro-, side, referring to narrow blackish lateral band from gill cover to base of caudal fin

Enteromius taeniurus (Boulenger 1903)     taenio-, band; oura, tail, referring to black band on each side of caudal peduncle

Enteromius tanapelagius (de Graaf, Dejen, Sibbing & Osse 2000)    tana, referring to Lake Tana, Ethiopia, where barb is part of an endemic cyprinid species flock; pelagius, pelagic, referring to occurrence in deeper, offshore waters

Enteromius tangandensis (Jubb 1954)    ensis, suffix denoting place: Tanganda River, Rhodesia, type locality

Enteromius tegulifer (Fowler 1936)    tegula, roofing tile; fero, to bear, referring to dark spots on sides “appearing like tiles”

Enteromius tetrastigma (Boulenger 1913)    tetra-, four; stigma, mark, referring to four rounded black spots on each side of body, two each above and below lateral line

Enteromius teugelsi (Bamba, Vreven & Snoeks 2011)    in honor of Guy G. Teugels (1954-2003), curator of fishes at the Musée Royale de l’Afrique Centrale, “an outstanding ichthyologist who introduced the first and second author to fish taxonomy and greatly contributed to the knowledge of the African fishes over the last twenty years”

Enteromius thamalakanensis (Fowler 1935)    ensis, suffix denoting place: Thamalakane River, Botswana, type locality

Enteromius thysi (Trewavas 1974)    in honor of ichthyologist D. Thys van den Audenaerde (b. 1934), who collected this species in Fernando Poo (Cameroon) and recorded difference in color pattern between it and E. trispilos

Enteromius tiekoroi (Lévêque, Teugels & Thys van den Audenaerde 1987)    in honor of fisherman Tiekoro Sineogo, with whom the authors have worked since 1975

Enteromius tomiensis (Fowler 1936)    ensis, suffix denoting place: Tomi River at Fort Sibut, Ubangi-Shari (now Central African Republic), type locality

Enteromius tongaensis (Rendahl 1935)    ensis, suffix denoting place: Tonga, in the Sudan, type locality

Enteromius toppini (Boulenger 1916)    in honor of F. Toppin, Natal Museum (South Africa), who collected type

Enteromius trachypterus (Boulenger 1915)    trachys, rough; pterus, fin, presumably referring to breeding tubercles on 2nd, 3rd and 4th branched rays of anal fin

Enteromius traorei (Lévêque, Teugels & Thys van den Audenaerde 1987)    in honor of friend and colleague Kassoum Traoré, l’Institut d’Écologie tropicale d’Abidjan, Ivory Coast, where it is endemic

Enteromius treurensis (Groenewald 1958)   ensis, suffix denoting place: Treur River, Transvaal, South Africa, type locality

Enteromius trimaculatus (Peters 1852)    tri-, three; maculatus, spotted, referring to three black spots, two on side and one at base of caudal peduncle

Enteromius trinotatus (Fowler 1936)    tri-, three; notatus, marked, referring to 3-5 black spots on sides

Enteromius trispiloides (Lévêque, Teugels & Thys van den Audenaerde 1987)    oides, having the form: E. trispilos, both of which have three round spots along middle of body

Enteromius trispilomimus (Boulenger 1907)    tri-, three; spilos, spot; mimus, imitator, referring to how three large round black spots on each side of body “strikingly resembles” those of E. trispilos

Enteromius trispilopleura (Boulenger 1902)    tri-, three; spilos, spot; pleura, side, referring to three round black spots on each side of body

Enteromius trispilos (Bleeker 1863)    tri-, three; spilos, spot, referring to three rounds spots along middle of body

Enteromius turkanae (Hopson & Hopson 1982)    of Lake Turkana, Kenya, where it is endemic

Enteromius unitaeniatus (Günther 1866)    uni-, one; taeniata, banded, referring to greyish or blackish band that runs from eye to caudal fin

Enteromius urostigma (Boulenger 1917)    oura-, tail; stigma, mark or spot, referring to round black spot at base of caudal fin

Enteromius usambarae (Lönnberg 1907)    of Usambara, presumably referring to region near Usambara Mountains in northeast Tanzania, near or overlooking type locality in Tanga

Enteromius validus (Stiassny, Liyandja & Monsembula Iyaba 2016)    strong or powerful, referring to its robust appearance

Enteromius vanderysti (Poll 1945)    in honor of entomologist R. P. Vanderyst, who collected type

Enteromius vandewallei Lederoun & Vreven 2016    in honor of Pierre Vandewalle, University of Liège (Belgium), who introduced the first author to fish taxonomy

Enteromius venustus (Bailey 1980)    pretty, referring to “attractive appearance of this small fish in life” (orange-red upper flanks, caudal peduncle and tail)

Enteromius viktorianus (Lohberger 1929)    anus, belonging to: Lake Victoria basin, Kenya, only known locality

Enteromius viviparus (Weber 1897)    livebearer, reflecting Weber’s mistaken belief that it is viviparous (he described unborn fry with yolk sacs in the ovary of his specimen, which were actually undigested fry from a cichlid species in the stomach)

Enteromius walkeri (Boulenger 1904)    in honor of the late Robert Bruce Napoleon Walker (1832-1901), West African trader, explorer, anthropologist and natural history collector, who collected type

Enteromius wellmani (Boulenger 1911)    in honor of tropical medicine physician F. C. Wellman, who collected type

Enteromius yeiensis (Johnsen 1926)    ensis, suffix denoting place: Yei River, Sudan, type locality

Enteromius yongei (Whitehead 1960)    in honor of zoologist Charles Maurice Yonge (1899-1986), for the “interest he has shown and the assistance he has given to many aspects of fishery research in East Africa”

Enteromius zalbiensis (Blache & Miton 1960)    ensis, suffix denoting place: Zalbi, Chad, paratype locality

Enteromius zanzibaricus (Peters 1868)    icus, belonging to: Zanzibar, a possible misnomer since the barb does not occur there; perhaps name refers to general vicinity of type locality (Mombassa, Kenya) to the Zanzibar Archipelago

Epalzeorhynchos Bleeker 1855    rhynchos, snout; epalzeo-, meaning unknown, but based on Bleeker’s habit of providing Dutch transliterations of Latin epithets in his 1860 Ichthyologiae Archipelagi Indici Prodromus, it means “horn” and refers to cone-shaped, cartilaginous protuberance on snout of E. kalopterum that can horizontally move nearly 90˚ away from snout, thus giving head a horned appearance [some online references say epalzeo– means “curative” but do not explain this interpretation]; ichthys, fish

Epalzeorhynchos bicolor (Smith 1931)    bi-, two, i.e., two-colored, reflecting how “rich velvety black of the body and most of the fins is strikingly relieved by the bright orange color of the caudal and pectoral fins, making this one of the most beautiful of the many attractively colored cyprinoid fishes of Siam”

Epalzeorhynchos frenatum (Fowler 1934)    brindled, referring to conspicuous black “bridle-like” band from snout to eye (though not crossing tip of snout)

Epalzeorhynchos kalopterum (Bleeker 1851)    kallos, beauty; pterus, fin, referring to pink or red fins, which Bleeker in 1860 described as pretty (“Fraaivinnige”)

Epalzeorhynchos munense (Smith 1934)    ensis, suffix denoting place: Menam Mun (or simply Mun) River, eastern Thailand, type locality